Hostname: page-component-848d4c4894-ttngx Total loading time: 0 Render date: 2024-06-07T07:44:38.839Z Has data issue: false hasContentIssue false

A Guadalupian-Lopingian (Middle to Late Permian) brachiopod fauna from the Juripu Formation in the Yarlung-Zangbo suture zone, southern Tibet, China

Published online by Cambridge University Press:  20 May 2016

G. R. Shi
Affiliation:
1School of Ecology and Environment, Deakin University, Melbourne Campus, 221 Burwood Highway, Burwood, Victoria 3125, Australia,
Shuzhong Shen
Affiliation:
2Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 Eastern Beijing Road, Nanjing, Jiangsu Province 210008, P. R. China,
Lipei Zhan
Affiliation:
3Institute of Geology, Chinese Academy of Geological Sciences, Baiwanzhuang Road, Beijing 100037, P. R. China

Abstract

A brachiopod fauna of 16 species belonging to 11 genera and three genera and species indeterminate from the middle and upper parts of the Juripu Formation in the Yarlung-Zangbo (Indus-Tsangbo) Suture zone (=Yarlung-Zangbo River zone), southern Tibet, is described and figured for the first time. A new species, Taeniothaerus zhongbaensis, is described. The fauna is comparable with that in the Kalabagh Member of the Wargal Formation of the Salt Range, Pakistan, and is considered to be most likely Capitanian (late Guadalupian/Middle Permian) to Wuchiapingian (early Lopingian/early Late Permian) in age, as indicated by the majority of the brachiopod species and by being constrained by an underlying fusulinacean fauna (Parafusulina Zone) and an overlying ammonite fauna (Cyclolobus fauna).

Type
Research Article
Copyright
Copyright © The Paleontological Society 

Access options

Get access to the full version of this content by using one of the access options below. (Log in options will check for institutional or personal access. Content may require purchase if you do not have access.)

References

Angiolini, L. 2001. Permian brachiopods from Karakorum (Pakistan). Rivista Italiana di Paleontologia e Stratigrafia, 107:307344.Google Scholar
Archbold, N. W. 1981. Quinquenella (Chonetidina, Brachiopoda) from the Permian of Western Australia. Journal of Paleontology, 55:204210.Google Scholar
Archbold, N. W. 1997. Studies on Western Australian Permian brachiopods 14. The fauna of the Artinskian High Cliff Sandstone, Perth Basin. Proceedings of the Royal Society of Victoria, 109:199231.Google Scholar
Archbold, N. W., and Thomas, G. A. 1986. Neospirifer and Trigonotreta (Spiriferida, Brachiopoda) from the Permian of Western Australia. Alcheringa, 10(2):125161.Google Scholar
Broili, F. 1916. Die Permischen brachiopoden von Timor. In Wanner, J. (ed.), Palaeontologie von Timor, 7, Stuttgart, 104 p.Google Scholar
Brunton, C. H. C., Lazarev, S. S., Grant, R. E., and Yugan, Jin. 2000. Productida, p. 424609. In Kaesler, R. (ed.), Treatise on Invertebrate Paleontology, Pt. H, Brachiopoda Revised, Volume 3. Geological Society of America and University of Kansas Press, New York and Lawrence.Google Scholar
Carter, J. L., Johnson, J. G., Gourvennec, R., and Hongfei, Hou. 1994. A revised classification of the spiriferid brachiopods. Annals of Carnegie Museum, 68:327374.Google Scholar
Chao, Y. T. 1927. Productidae of China, Pt. 1, Producti. Palaeontologica Sinica, series B, volume 5, Fascicle 2:1133.Google Scholar
Conrad, T. A. 1839. Second annual report on the palaeontological department of the survey. New York Geological Survey, Annual Report, 3:5766.Google Scholar
Davidson, T. 1862. On some Carboniferous Brachiopoda collected in India by A. Fleming, M.D. and W. Purdon, Esq., F.G.S. Quarterly Journal of the Geological Society of London, 18:2535.Google Scholar
Diener, C. 1897. Himalayan fossils. The Permocarboniferous fauna of Chitichun, No.1. Geological Survey of India, Memoirs, Palaeontologia Indica, Series 15, 1(3):1105.Google Scholar
Diener, C. 1899. Himalayan fossils. Anthracolithic fossils of Kashmir and Spiti. Memoirs of the Geological Survey of India, Palaeontologia Indica, Series 15, 1(2):195.Google Scholar
Diener, C. 1903. Permian fossils of the Central Himalayas. Memoirs of the Geological Survey of India, Palaeontologia Indica, Series 15, 1(5):1204.Google Scholar
Diener, C. 1915. The Anthracolithic faunae of Kashmir, Kanaur and Spiti. Memoirs of the Geological Survey of India, Palaeontologia Indica, New Series 5, 2:1135.Google Scholar
Ding (Ting) Peichen. 1962. Some Upper Permian brachiopods from Tibet. Acta Palaeontologica Sinica, 10(4):451471. (In Chinese)Google Scholar
Rensen, Fang, and Jiancai, Fan. 1994. Middle to Upper Carboniferous-Early Permian Gondwana facies and palaeontology in Western Yunana. Yunnan Science and Technology Press, Kunming, 121 p.Google Scholar
Ruling, Feng, and Zongrong, Jiang. 1978. Brachiopoda, p. 231305. In Guizhou Working Group of Stratigraphy and Paleontology (ed.), Paleontological Atlas of Southwest China. Guizhou Volume. Pt. 2 (Carboniferous to Quaternary). Geological Publishing House, Beijing. (In Chinese)Google Scholar
de Waldheim, G. Fischer 1825. Notice sur la Choristite, genre de Coquilles bivalves fossiles du gouvernement de Moscou. Programme d'invitation a la Societe Imperiale des Naturalistes de Moscou, 11 p.Google Scholar
Frech, F. 1911. Das Obercarbon Chinas, Die Dyas, p. 1289. In von Richthofen, F. (ed.), China, Volume 5.Dietrich Reimer, Berlin.Google Scholar
Fredericks, G. N. 1924. Paleontologicheskie etiudy. 2. O verkhne-kamennougol'nykh spiriferidakh Urala [Paleontological studies. 2. On some Upper Carboniferous spiriferids of the Urals]. Izvestiia Geologicheskogo Komiteta, 38(3):295324. (In Russian)Google Scholar
Garzanti, E., Angiolini, L., and Sciunnach, D. 1996. The Permian Kuling Group (Spiti, Lahaul and Zanskar; NW Himalaya): sedimentary evolution during rift/drift transition and initial opening of Neo-Tethys. Rivista Italiana di Paleontologia e Stratigrafa, 102(2):175200.Google Scholar
Geological Survey of Tibet. 1983. 1:1000000 geological map of the Xigaze (H–45) and Yadong (G–45) Districts (Xizang) and explanatory notes. Bureau of Geology and Mineral Resources of the Xizang Autonomous Region, Lhasa, 568 p.Google Scholar
Grabau, A. W. 1934. Early Permian fossils of China, I. Early Permian brachiopods, pelecypods and gastropods of Kueichow. Palaeontologica Sinica, Series B, 8(3):1168.Google Scholar
Grant, R. E. 1965. The brachiopod Superfamily Stenoscismatacea. Smithsonian Miscellaneous Collections, 148(2):1192.Google Scholar
Grant, R. E. 1976. Permian brachiopods from Southern Thailand. The Paleontological Society Memoir 9 [(supplement to Journal of Paleontology, 50(3)], 269 p.Google Scholar
Tieying, Guo, Dingyi, Liang, Yizhi, Zhang, and Chonghe, Zhao. 1991. Geology of Ngari, Tibet (Xizang). China University of Geosciences Press, Wuhan, 464 p. (In Chinese)Google Scholar
Hamlet, B. 1928. Permische Brachiopoden, Lamellibranchiaten und Gastropoden von Timor. Jaarboek van het Mijnwezen in Nederlandsch Oost-Indie, Gravenhage, 56(2):1109.Google Scholar
Hayasaka, I., and Gan, S. 1940. A note on Camarophoria ‘purdoni’ from the Permian of Timor. Journal of the Geological Society of Japan, 47(558):127132.Google Scholar
Hongfei, Hou, Lipei, Zhan, and Binwei, Chen. 1979. The Coal-bearing Strata and Fossils of the Late Permian from Guangdong. Geological Publishing House, Beijing, 116 p. (In Chinese)Google Scholar
Huang, T. K. 1932. Late Permian brachiopod fossils of southwestern China. Palaeontologica Sinica, Series B, 9(1):1138.Google Scholar
Jianjun, Jiang, and Haijun, Yan. 1992. On the characteristics, members, geological age and geographical distribution of Stereochia and Costiferina (Brachiopoda). Journal of China University of Geosciences, 6(2):174182. (In Chinese)Google Scholar
Yugan, Jin, and Shizhong, Hu. 1978. Brachiopods of the Kuhfeng Formation in South Anhui and Nanking Hills. Acta Palaeontologica Sinica, 17(2):101127.Google Scholar
King, W. 1865. Remarks on the histology of two specimens of Rhynchopora geinitziana de Verneuil, from near the River Oukhla, Province of Archangel, and St. Petersburg. Annals and Magazine of Natural History, series 3, 16:124128.Google Scholar
Kotlyar, G. V. 1964. Novye predstaviteli roda Productus iz verkhnei permi iuzhnogo Primor'ia [New representative of the genus Productus from the Upper Permian of South Primorye]. Vsesoiuznyi Nauchno-Issledovatel'skii Geologicheskii Institut (VSEGEI), Trudy, 93:123134.Google Scholar
Legrand-Blain, M. 1977 (1976). Repartition du groupe de Spiriferella rajah (Salter, 1865) et description de Spiriferella nepalensis, nov. sp., appartenant a ce groupe. Centre National de la Recherche Scientifique, Colloques Internationaux, France, 268(2):237250.Google Scholar
Leven, E. Ya. 1993. Main events in Permian history of the Tethys and fusulinids. Stratigraphy and Geological Correlation, 1(1):5165.Google Scholar
Leven, E. Ya., Grunt, T. S., Jing-dang, Lin, and Liang-fang, Li. 2001. Upper Permian stratigraphy of the Zhesi Honguer area (North China). Stratigraphy and Geological Correlation, 9(5):441453.Google Scholar
Zhuoting, Liao. 1980. Upper Permian brachiopods from western Guizhou, p. 241277. In Nanjing Institute of Geology and Paleontology (ed.), Stratigraphy and Paleontology of the Upper Permian Coal-bearing Formation in Western Guizhou and Eastern Yunnan. Science Press, Beijing. (In Chinese)Google Scholar
(=Licharew), B. K. Likharev 1937 (1936). Permskie brachiopody Severnogo Kavkaza. Semeustva Chonetidae Hall et Clarke i Productidae Gray [Brachiopoda of the Permian System of the USSR. Fascicule 1. Permian Brachiopoda of North Caucasus: Families Chonetidae Hall & Clarke and Productidae Gray] Monografii po Palaeontologii SSSR, 39, 1152. (In Russian)Google Scholar
(=Licharew), B. K. Likharev, and Kotlyar, G. V. 1978. Permskie brakhiopody iuzhnogo Primor'ia [Permian brachiopods of South Primorye], p. 6375, 96–99. In Popeko, L. I. (ed.), Verkhnii Paleozoi Severo-Vostochnoi Azii [Upper Paleozoic of Northeast Asia]. Akademiya Nauk SSSR Dal'nevostochnyie Nauchnyie Tsentr Institut Tektoniki i Geofiziki, Vladivostok.Google Scholar
McCoy, F. 1844. A Synopsis of the Characters of the Carboniferous Limestone Fossils of Ireland. Williams & Norgate, London, 207 p.Google Scholar
Morris, J. 1845. Account of the fossil Mollusca, p. 270291. In de Strzelecki, P. E. (ed.), Physical Description of New South Wales and Van Diemen's Land. Longman, Brown, Green and Longmans, London.Google Scholar
Muir-Wood, H. M. 1955. A history of the classification of the phylum Brachiopoda. British Museum (Natural History), Monograph, viii + 124 p.Google Scholar
Muir-Wood, H. M. 1962. On the morphology and classification of the brachiopod Suborder Chonetoidea. British Museum (Natural History), Monograph, viii + 132 p.Google Scholar
Muir-Wood, H. M., and Cooper, G. A. 1960. Morphology, classification and life habits of the Productoidea (Brachiopoda). Memoir of the Geological Society of America, 81:1447.CrossRefGoogle Scholar
Muir-Wood, H. M., and Oakley, K. P. 1941. Upper Palaeozoic faunas of North Sikkim. Memoirs of Geological Survey of India, Palaeontologia Indica, 31(1):191.Google Scholar
Oehlert, D. P. 1887. Appendice sur les brachiopodes, p. 11891334. In Fischer, P. (ed.), Manuel de conchyliologie et de paleontologie conchyliologue, ou histoire naturelle des mollusques vivants et fossiles, Vol. 4, part 11. F. Savy, Paris.Google Scholar
Parfrey, S. M. 1983. Redescription of the Permian aulostegid brachiopod Taeniothaerus subquadratus (Morris, 1845), the type species of Taeniothaerus Whitehouse. Memoir of the Association of Australasian Palaeontologists, 1:289296.Google Scholar
Plodowski, G. 1968. Neue spiriferen aus Afghanistan. Senckenbergiana Lethaea, 49(2–3):251258.Google Scholar
Racheboeuf, P. R. 2000. Chonetidina, p. 362423. In Kaesler, R. (ed.), Treatise on Invertebrate Paleontology, Pt. H, Brachiopoda Revised, Volume 2. Geological Society of America and University of Kansas Press, New York and Lawrence.Google Scholar
Reed, F. R. C. 1944. Brachiopoda and Mollusca from the Productus Limestones of the Salt Range. Memoirs of the Geological Survey of India, Palaeontologia Indica, 23(2):1678.Google Scholar
Ruzhentsev, V. E., and Sarytcheva, T. G. 1965. Razivitie i smena morskikh organizmov na rubezhe Paleozoia i Mezozoia [The development and change of marine organisms at the Paleozoic-Mesozoic boundary]. Akademiia Nauk SSSR, Paleontologicheskii Institut, Trudy, 108:1370. (In Russian)Google Scholar
Sarytcheva, T. G., Likharev, B. K., and Sokolskaja, A. N. 1960. Otriad Productida, p. 221238. In Orlov, Y. A. (ed.), Osnovy Paleontologii; Mshanki Brakhiopody. Izdatel'stvo Akademii Nauk SSSR, Moscow. (In Russian)Google Scholar
Savage, N. M., Mancenido, M. O., and Owen, E. F. 2002. Order Rhynchonellida, p. 10271376. In Kaesler, R. (ed.), Treatise on Invertebrate Paleontology, Pt. H, Brachiopoda Revised, Volume 4. Geological Society of America and University of Kansas Press, New York and Lawrence.Google Scholar
Schuchert, C. 1893. A Classification of the Brachiopoda. American Geologist, 11(3):141167.Google Scholar
Schuchert, C. 1929. Classification of brachiopod genera, fossil and recent, p. 1025. In Schuchert, C. and LeVene, C. M., Animalia pars 42 [In Pompeckj, J.F. (ed.), Fossilium Catalogus, Volume 1] Junk, Berlin.Google Scholar
Shuzhong, Shen, and Yugan, Jin. 1999. Brachiopods from the Permian-Triassic boundary beds at the Selong Xishan section, Xizang (Tibet), China. Journal of Asian Earth Sciences, 17(4):547559.Google Scholar
Shuzhong, Shen, Archbold, N. W., Shi, G. R., and Chen, Z. Q. 2000. Permian brachiopods from the Selong Xishan section, Xizang (Tibet), China, Pt. 1, Stratigraphy, Strophomenida, Productida and Rhynchonellida. Geobios, 33:725752.Google Scholar
Shuzhong, Shen, Archbold, N. W., Shi, G. R., and Chen, Z. Q. 2001a. Permian brachiopods from the Selong Xishan section, Xizang (Tibet), China, Pt. 2, Palaeobiogeographical and palaeoecological implications, Spiriferida, Athyridida and Terebratulida. Geobios, 34:157182.Google Scholar
Shuzhong, Shen, Archbold, N. W., and Shi, G. R. 2001b. A Lopingian (Late Permian) brachiopod fauna from the Qubuerga Formation at Shengmi in the Mount Qomolangma region of Southern Xizang (Tibet), China. Journal of Paleontology, 75:274283.Google Scholar
Shuzhong, Shen, Changqun, Cao, Shi, G. R., Xiangdong, Wang, Shilong, Mei, and Yugan, Jin. 2003. Lopingian (Late Permian) stratigraphy and deposition in southern Tibet and its correlation over the peri-Gondwana region. Newsletter of Stratigraphy, 39:157179.Google Scholar
Shuzhong, Shen, Shi, G. R., and Archbold, N. W. 2003a. A Wuchiapingian (Late Permian) brachiopod fauna from an exotic block in the Indus-Tsangpo Suture zone, southern Tibet and its palaeobiogeographical and tectonic implications. Palaeontology, 46:225256.Google Scholar
Shuzhong, Shen, Shi, G. R., and Archbold, N. W. 2003b. Lopingian (Late Permian) brachiopods from the Qubuerga Formation at the Qubu section in the Mt. Qomolongma region, southern Tibet (Xizang), China. Palaeontographica Abt. A, 268:49101.Google Scholar
Shuzhong, Shen, Dongli, Sun, and Shi, G. R. 2003c. A Capitanian (late Middle Permian) brachiopod fauna from an exotic limestone block at Xiukang in Lhaze County of Tibet. Journal of Asian Earth Sciences, 21:11251137.Google Scholar
Huaibin, Sheng. 1989. Early Permian ammonoids from the Juripu Formation, Zhonga, Xizang (Tibet). Geological Review, 35:324332. (In Chinese)Google Scholar
Shi, G. R., and Shuzhong, Shen. 1997. A Late Permian brachiopod fauna from Selong, Southern Xizang (Tibet), China. Proceedings of the Royal Society of Victoria, 109(1):3756.Google Scholar
Shimizu, D. 1981. Upper Permian brachiopod fossils from Guryul Ravine and its spur three kilometers of Barus. Palaeontologica Indica, New Series, 46:6785.Google Scholar
Stehli, F. G. 1954. Lower Leonardian Brachiopoda of the Sierra Diablo. Bulletin of the American Museum of Natural History, 105(3):257358.Google Scholar
Zhengxiang, Tong. 1978. Brachiopoda, p. 210267. In Geological Institute of Southwest China (ed.), Paleontological Atlas of Southwest China, Sichuan Province, Volume 2 (Carboniferous and Permian). Geological Publishing House, Beijing. (In Chinese)Google Scholar
Tschernyschev, T. N. 1902. Verkhnekamennougol'nye brakhiopody Urala i Timana [Upper Carboniferous brachiopods from the Urals and Timan]. Trudy Geologicheskogo Komiteta, 16(2):1749. (In Russian)Google Scholar
Waagen, W. 1883–1884. Salt Range fossils I. Productus-Limestone fossils. Geological Survey of India, Memoirs, Palaeontologia Indica, Series 13, facicules 2– 4, 391728, pls. 29–81.Google Scholar
Chengshan, Wang, Daixiang, Xia, Xiang, Zhou, and Jianping, Chen. 1999. Geology between the Indus-Yarlung Zangbo Suture Zone and the Himalaya Mountains, Xizang (Tibet), China. Geological Publishing House, Beijing, 160 p. (In Chinese)Google Scholar
Guoping, Wang, Qingzhao, Liu, Yugan, Jin, Shizhong, Hu, Wenping, Liang, and Zhuoting, Liao. 1982. Brachiopoda, p. 186256. In Institute of Geology and Mineral Resources (ed.), Paleontological Atlas of East China. 2. Late Paleozoic. Geolological Publishing House, Beijing. (In Chinese)Google Scholar
Yu, Wang, Yugan, Jin (Yu-kan, Ching), and Dawei, Fang. 1964. Brachiopod Fossils of China. Science Press, Beijing, Volumes 1 and 2, 777 p. (In Chinese)Google Scholar
Yujing, Wang, Jinzhang, Sheng, and Linxin, Zhang. 1981. Fusulinids from Xizang of China, p. 180. In Chinese Academy of Sciences Integrated Scientific Expedition Team to the Qinghai-Xizang (Tibet) Plateau (ed.), The Series of the Scientific Expedition to the Qinghai-Xizang Plateau. Palaeontology of Xizang, Book III. Science Press, Beijing. (In Chinese)Google Scholar
Wardlaw, B. R., and Pogue, K. R. 1995. The Permian of Pakistan, p. 215224. In Scholle, P. A., Peryt, T. M., and Ulmer-Scholle, D. S. (eds.), The Permian of Northern Pangea 2, Sedimentary Basins and Economic Resources. Springer-Verlag, Berlin.Google Scholar
Wardlaw, B. R., and Shi-long, Mei. 1999. Refined conodont biostratigraphy of the Permian and lowest Triassic of the Salt and Khizor Ranges, Pakistan, p. 154156. In Hongfu, Yin and Iinnan, Tong (eds.), Proceedings of the International Conference on Pangea and the Paleozoic-Mesozoic Transition. China University of Geosciences Press, Wuhan, China.Google Scholar
Waterhouse, J. B. 1966. Lower Carboniferous and Upper Permian brachiopods from Nepal. Geologische Bundesanstalt, Jahrbuch, 12:599.Google Scholar
Waterhouse, J. B. 1968. The classification and descriptions of Permian Spiriferida (Brachiopoda) from New Zealand. Palaeontographica, Abt. A, 129(1–3):194.Google Scholar
Waterhouse, J. B. 1970. Permian brachiopod Retimarginifera gen. nov. from the Byro Group of Carnarvon Basin, Western Australia. Proceedings of the Royal Society of Western Australia, 53(4):120128.Google Scholar
Waterhouse, J. B. 1975. New Permian and Triassic brachiopod taxa. Papers of Department of Geology, University of Queensland, 7(1):123.Google Scholar
Waterhouse, J. B. 1978. Permian Brachiopoda and Mollusca from North-West Nepal. Palaeontographica, Abt. A, 160(1–6):1175.Google Scholar
Waterhouse, J. B. 1983. Permian brachiopods from Pija Member, Senja Formation in Manang district of Nepal, with new brachiopod genera and species from other regions. Indian Geologists' Association Bulletin, 16(2):111151.Google Scholar
Waterhouse, J. B. 1987. Late Palaeozoic Brachiopoda (Athyrida, Spiriferida and Terebratulida) from the southeast Bowen Basin, east Australia. Palaeontographica Abt., 196:156.Google Scholar
Whitehouse, F. W. 1928. Notes on Upper Palaeozoic marine horizons in eastern and western Australia. Australasian Association for the Advancement of Science, Report, 18:281283.Google Scholar
Williams, A., and Harper, D. A. 2000. Orthida, p. 714844. In Kaesler, R. (ed.), Treatise on Invertebrate Paleontology, Pt. H, Brachiopoda Revised, Volume 3. Geological Society of America and University of Kansas Press, New York and Lawrence.Google Scholar
Deli, Yang, Shizhao, Ni, Meili, Chang, and Ruxuan, Zhao. 1977. Brachiopoda, p. 303470. In Hubei Geological Institute, and Geological Bureaus of Henan, Hubei, Hunan, Guangdong, and Guangxi Provinces (eds.), Paleontological Atlas of Central South China, Volume 2 (Late Paleozoic). Geological Publishing House, Beijing. (In Chinese)Google Scholar
Zunyi, Yang, and Zetong, Nie. 1990. Palaeontology of Nagri, Tibet (Xizang). China University of Geosciences Press, Wuhan, China, 380 p.Google Scholar
Zunyi, Yang, and Kangfu, Zhang. 1982. Fossils in the upper part of the Lower Permian from Surishan, Dingri, Xizang, p. 310315. In Monograph on Mount Xixia Bangma Scientific Expedition 1964, Science Press, Beijing. (In Chinese)Google Scholar
Jixiang, Yin. 1997. Stratigraphic Geology of Gondwana Facies of Qinghai-Xizang (Tibet) Plateau and Adjacent Areas. Geological Publishing House, Beijing, 205 p. (In Chinese)Google Scholar
Lipei, Zhan, and Rangyong, Wu. 1982. Early Permian brachiopods from Xainza district, Xizang (Tibet), p. 86109. In Contribution to the Geology of the Qinghai-Xizang (Tibet) Plateau, 7. Geological Publishing House, Beijing.Google Scholar
Shouxin, Zhang (Chang), and Yugan, Jin (Ching). 1976. Late Paleozoic brachiopods from the Mount Jolmo Lungma region, p. 159271. In Xizang Scientific expedition team of Chinese Academy of Sciences (ed.), A Report of Scientific Expedition in the Mount Jolmo Lungma Region (1966–1968), 2, Science Press, Beijing. (In Chinese)Google Scholar
Yan, Zhang (Chang), and Yugan, Jin (Ching). 1961. Late Permian brachiopod fossils from Jin Xian, Anhui province. Acata Palaeontologica Sinica, 9(4):401414.Google Scholar