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The Late Miocene Talpidae (Eulipotyphla, Mammalia) from the Pannonian Region, Slovakia

Published online by Cambridge University Press:  03 April 2024

Florentin Cailleux*
Affiliation:
Comenius University in Bratislava, Faculty of Natural Sciences, Department of Geology and Paleontology, Ilkovičova 6, Mlynská dolina G, SK–842 15 Bratislava, Slovakia. , Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, The Netherlands.
Lars W. van den Hoek Ostende
Affiliation:
Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, The Netherlands.
Peter Joniak
Affiliation:
Comenius University in Bratislava, Faculty of Natural Sciences, Department of Geology and Paleontology, Ilkovičova 6, Mlynská dolina G, SK–842 15 Bratislava, Slovakia. ,
*
*Corresponding author.

Abstract

Central Europe is an area of high diversity for the Talpidae (Eulipotyphla, Mammalia) during the Late Miocene. The assemblages from Slovakia (Borský Svätý Jur, Krásno, Pezinok, Šalgovce, Studienka, Triblavina) are no exception with their abundant material representing eleven species. The uropsiline Desmanella is represented by D. rietscheli and D. dubia. Desmanini fossils are attributed to Archaeodesmana vinea, Archaeodesmana dissona new species, Gerhardstorchia biradicata, and Gerhardstorchia sp. The scalopines Proscapanus minor and P. austriacus are well recorded in the Vallesian localities and support the emergence of P. austriacus before the MN9/10 transition. Talpini and Urotrichini are especially rare and only represented by Talpa cf. T. minuta and Urotrichini gen. sp. indet. Finally, we identified the youngest occurrence of Desmanodon in Europe, D. cf. D. fluegeli, at the MN9 locality of Borský Svätý Jur. The high diversity in the Late Miocene Central European is partly explained by the co-occurrence of the competing Scalopini and Talpini during the Vallesian, indicating high resource environments. The decline of these tribes, followed by the success of the desmans during the Turolian, appears as a consequence of regional environmental changes.

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Copyright © The Author(s), 2024. Published by Cambridge University Press on behalf of Paleontological Society

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References

Bachmayer, F., and Wilson, R.W., 1978, A second contribution to the fossil small mammal fauna of Kohfidisch, Austria: Annalen des Naturhistorischen Museums in Wien, v. 81, p. 129161.Google Scholar
Bannikova, A., Zemlemerova, E.D., Colangelo, P., Sözen, M., Sevindik, M., Kidov, A.A., Dzuev, R.I., Kryštufek, B., and Lebedev, V.S., 2015, An underground burst of diversity—a new look at the phylogeny and taxonomy of the genus Talpa Linnaeus, 1758 (Mammalia: Talpidae) as revealed by nuclear and mitochondrial genes: Zoological Journal of the Linnean Society, v. 175, p. 930948, https://doi.org/10.1111/zoj.12298.CrossRefGoogle Scholar
Bilgin, M., Joniak, P., Mayda, S., Göktaş, F., Kaya, T., Campomanes, P.P., and van den Hoek Ostende, L.W., 2019, Sabuncubeli too, Bornova, a second micromammal assemblage from the Sabuncubeli section (Early Miocene, western Anatolia): Palaeobiodiversity and Palaeoenvironments, v. 99, p. 655671, https://doi.org/10.1007/s12549-019-00395-2.CrossRefGoogle Scholar
Cailleux, F., Joniak, P., and van den Hoek Ostende, L.W., 2023, The Late Miocene Erinaceidae (Eulipotyphla, Mammalia) from the Pannonian Region, Slovakia: Journal of Paleontology, v. 97, p. 777798, https://doi.org/10.1017/jpa.2023.50.CrossRefGoogle Scholar
Crochet, J.-Y., 1986, Insectivores Pliocènes du sud de la France (Languedoc–Roussillon) et du nord-est de l'Espagne: Palaeovertebrata, v. 16, p. 145171.Google Scholar
Crochet, J.-Y., and Green, M., 1982, Contributions à l'étude des micromammifères du gisement Miocène Supérieur de Montredon (Hérault). Les insectivores: Palaeovertebrata, v. 12, p. 119131.Google Scholar
Dahlmann, T., 2001, Die Kleinsäuger der unter-pliozänen Fundstelle Wölfersheim in der Wetterau (Mammalia: Lipotyphla, Chiroptera, Rodentia): Courier Forschungsinstitut Senckenberg, v. 227, p. 1129.Google Scholar
Dahlmann, T., and Doğan, S., 2011, Gerhardstorchia nomen novum: a new name for Storchia Dahlmann 2001 (Mammalia: Lipotyphla: Talpidae: Desmaninae): Paläontologische Zeitschrift, v. 85, p. 9191, https://doi.org/10.1007/s12542-010-0079-4.CrossRefGoogle Scholar
de Blainville, H.M.D., 1840, Osteographie des mammifères insectivores (Talpa, Sorex et Erinaceus L.): Osteographie des Mammifères, v. 1, p. 1115.Google Scholar
Dobson, G.E., 1883, A Monograph of the Insectivora, Systematic and Anatomical: London, John van Voorst, v. 2, p. 86172.Google Scholar
Doukas, C.S., van den Hoek Ostende, L.W., Theocharopoulos, C.D., and Reumer, J.W.F., 1995, The vertebrate locality Maramena (Macedonia, Greece) at the Turolian–Ruscinian boundary (Neogene): Münchner Geowissenschaftliche Abhandlungen, v. 28, p. 4364.Google Scholar
Engesser, B., 1972, Die obermiozäne säugetierfauna von Anwil (Baselland) [Ph.D. thesis]: University of Basel, Liestal, Switzerland, Lüdin A.G., 363 p.Google Scholar
Engesser, B., 1980, Insectivora und Chiroptera (Mammalia) aus dem Neogen der Turkei: Schweizerische Paläontologische Abhandlungen, v. 102, p. 47159.Google Scholar
Engesser, B., 2009, The insectivores (Mammalia) from Sansan (Middle Miocene, south-western France): Schweizerische Paläontologische Abhandlungen, v. 128, p. 191.Google Scholar
Fejfar, O., and Sabol, M., 2005, Czech Republic and Slovak Republic, in van den Hoek Ostende, L.W., Doukas, C.S., and Reumer, J.W.F., eds., The Fossil Record of the Eurasian Neogene Insectivores (Erinaceomorpha, Soricomorpha, Mammalia), Part I: Scripta Geologica Special Issue, v. 5, p. 51–60.Google Scholar
Fischer, G., 1814, Zoognosia Tabulis Synopticis Illustrata, Volumen Tertium: Quadrupedum Reliquorum, Cetorum et Monotrymatum Descriptionem Continens: Moscow, Nicolai Sergeidis Vsevolozsky, 732 p.Google Scholar
Freudenberg, W., 1914. Die Säugetiere des Altern Quartärs von Mitteleuropa mit besonderes Berücksichtigung der Faunen von Hundsheim und Deutsch–Altenburg in Niederösterreich nebst Bemer-kungen über verwandte Formen anderer Fundorte: Geologische und Paläeontologische Abhandlungen, v. 12, p. 455670.Google Scholar
Furió, M., 2007, Los insectívoros (Soricomorpha, Erinaceomorpha, Mammalia) del Neógeno Superior del Levante Ibérico [Ph.D. thesis]: Barcelona, Spain, Universitat Autònoma de Barcelona, 299 p.Google Scholar
Furió, M., Van Dam, J., and Kaya, F., 2014, New insectivores (Lipotyphla, Mammalia) from the Late Miocene of the Sivas Basin, Central Anatolia: Bulletin of Geosciences, v. 89, p. 163181, https://doi.org/10.3140/bull.geosci.1472.Google Scholar
Gaillard, C., 1899, Mammifères Miocènes nouveaux ou peu connus de La Grive–Saint-Alban (Isère): Archives du Muséum d'Histoire Naturelle de Lyon, v. 7, 78 p.CrossRefGoogle Scholar
García-Alix, A., Furió, M., Minwer-Barakat, R., Martin Suarez, E., and Freudenthal, M., 2011, Environmental control on the biogeographical distribution of Desmanella (Soricomorpha, Mammalia) in the Miocene of the Iberian Peninsula: Palaeontology, v. 54, p. 753762.10.1111/j.1475-4983.2011.01062.xCrossRefGoogle Scholar
Gibert, J., 1975, New insectivores from the Miocene of Spain: Proceedings van de Koninklijke Nederlandse Akademie van Wetenschappen, v. 78, p. 108133.Google Scholar
Harrison, D.L., and Rzebik-Kowalska, B., 1994, A note on the occurrence of Desmanella cf. dubia Rümke, 1976 (Insectivora: Talpidae: Uropsilinae) in the Lower Pliocene of Podlesice, Poland: Cranium, v. 11, p. 36.Google Scholar
He, K., Shinohara, A., Helgen, K.M., Springer, M.S., Jiang, X.L., and Campbell, K.L., 2016, Talpid mole phylogeny unites shrew moles and illuminates overlooked cryptic species diversity: Molecular Biology and Evolution, v. 34, p. 7887, https://doi.org/10.1093/molbev/msw221.CrossRefGoogle ScholarPubMed
Hutchison, J.H., 1974, Notes on type specimens of European Miocene Talpidae and a tentative classification of old world Tertiary Talpidae (Insectivora: Mammalia): Geobios, v. 7, p. 211256.Google Scholar
Joniak, P., 2005, New rodent assemblages from the Upper Miocene deposits of the Vienna Basin and Danube Basin [thesis]: Bratislava, Slovakia, Department of Geology and Paleontology, Faculty of Natural Sciences, Comenius University in Bratislava, p. 1–126.Google Scholar
Joniak, P., 2016, Upper Miocene rodents from Pezinok in the Danube Basin, Slovakia: Acta Geologica Slovakia, v. 8, p. 114.Google Scholar
Joniak, P., and Šujan, M., 2020, Systematic and morphometric data of Late Miocene rodent assemblage from Triblavina (Danube Basin, Slovakia): Data in Brief, v. 28, e104961. https://doi.org/10.1016/j.dib.2019.104961.CrossRefGoogle ScholarPubMed
Joniak, P., Šujan, M., Fordinál, K., Braucher, R., Rybár, S., Kováčová, M., Kováč, M., and Aster Team, 2020, The age and paleoenvironment of a Late Miocene floodplain alongside Lake Pannon: rodent and mollusk biostratigraphy coupled with authigenic 10Be/9Be dating in the northern Danube Basin of Slovakia: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 538, e109482, https://doi.org/10.1016/j.palaeo.2019.109482.CrossRefGoogle Scholar
Kälin, D., and Engesser, B., 2001, Die jungmiozäne Säugetierfauna vom Nebelbergweg bei Nünningen (Kanton Solothurn, Schweiz): Schweizerische Paläontologische Abhandlungen, v. 121, p. 16.Google Scholar
Klietmann, J., 2013, Systematic and ecological analysis of Marsupialia and Eulipotyphla from Petersbuch 28 (Germany, Lower Miocene) [Ph.D. thesis]: Vienna, Austria, University of Vienna, 484 p.Google Scholar
Klietmann, J., Metscher, B.D., van den Hoek Ostende, L.W., Nagel, D., and Rummel, M., 2013, First record of an upper deciduous molar in Desmanella (Uropsilinae, Talpidae, Mammalia): Geobios, v. 46, p. 503510. https://doi.org/10.1016/j.geobios.2013.07.005.CrossRefGoogle Scholar
Klietmann, J., Nagel, D., Rummel, M., and van den Hoek Ostende, L.W., 2015, A gap in digging: the Talpidae of Petersbuch 28 (Germany, Early Miocene): Paläontologische Zeitschrift, v. 89, p. 563592. https://doi.org/10.1007/s12542-014-0228-2.CrossRefGoogle Scholar
Kormos, T., 1913, Trois nouvelles especes fossiles des Desmans en Hongrie: Annales Historico-naturales Musei Nationalis Hungarici, v. 11, p. 135145.Google Scholar
Kormos, T., 1930, Diagnosen neuer Säugetiere aus dem Oberpliozänen fauna des Somlyóberges bei Püspökfürdö: Annales Musei Nationalis Hungarici, v. 27, p. 237246.Google Scholar
Lartet, E., 1851, Notice sur la colline de Sansan: Annuaire du Département du Gers, 42 p.Google Scholar
Linnaeus, C., 1758, Systema Naturae per Regna Tria Naturae (tenth edition), Volume 1, Regnum Animale: Stockholm, Laurentii Salvii, 824 p.Google Scholar
Martín-Suárez, E., Bendala, N., and Freudenthal, M., 2001, Archaeodesmana baetica, sp. nov. (Mammalia, Insectivora, Talpidae) from the Mio–Pliocene transition of the Granada Basin, southern Spain: Journal of Vertebrate Paleontology, v. 21, p. 547554.CrossRefGoogle Scholar
Mayr, H., and Fahlbusch, V., 1975, Eine unterpliozäne Kleinsäugerfauna aus der Oberen Süßwasser–Molasse Bayerns: Mitteilungen der Bayerischen Staatssammlung für Paläontologie und Historische Geologie, v. 15, p. 91111.Google Scholar
Mein, P., 1999, The Late Miocene small mammal succession from France, with emphasis on the Rhône Valley localities, in Agustí, J., Rook, L., and Andrews, P., eds., Hominoid Evolution and Climatic Change in Europe, 1: Cambridge University Press, p 140164.CrossRefGoogle Scholar
Ménouret, B., and Mein, P., 2008, Les vertébrés du Miocène supérieur de Soblay (Ain, France): Travaux et Documents des Laboratoires de Géologie de Lyon, v. 165, p. 397.Google Scholar
Minwer-Barakat, R., García-Alix, A., and Freudenthal, M., 2008, Desmaninae (Talpidae, Mammalia) from the Pliocene of Tollo de Chiclana (Guadix Basin, Southern Spain): Considerations on the phylogeny of the genus Archaeodesmana. Geobios, v. 41, p. 381398. https://doi.org/10.1016/j.geobios.2007.08.001.Google Scholar
Minwer-Barakat, R., García-Alix, A., Martín-Suárez, E., and Freudenthal, M., 2020, Early Pliocene Desmaninae (Mammalia, Talpidae) from southern Spain and the origin of the genus Desmana: Journal of Vertebrate Paleontology, v. 40, n. e1835936, https://doi.org/10.1080/02724634.2020.1835936.CrossRefGoogle Scholar
Norris, R.W., and Kilpatrick, C.W., 2007, A high elevation record of the star-nosed mole (Condylura cristata) in northeastern Vermont: The Canadian Field-Naturalist, v. 121, p. 206207.10.22621/cfn.v121i2.447CrossRefGoogle Scholar
Oudemans, A.C., 1923, Acarologische Aanteekeningen. LXXX: Entomologische Berichten (Amsterdam), v. 6, p. 145155.Google Scholar
Petényi, S.J., 1864, Hátrahagyott munkái: Pest, MTA Kiadása, 130 p.Google Scholar
Piras, P., Sansalone, G., Teresi, L., Kotsakis, T., Colangelo, P., and Loy, A., 2012, Testing convergent and parallel adaptations in talpids humeral mechanical performance by means of geometric morphometrics and finite element analysis. Journal of Morphology, v. 273, p. 696711. https://doi.org/10.1002/jmor.20015.CrossRefGoogle ScholarPubMed
Prieto, J., 2010, The Middle Miocene mole Desmanodon crocheti sp. nov. (Talpidae, Mammalia): the last representative of the genus in the North Alpine foreland basin: Paläontologische Zeitschrift, v. 84, p. 217225, https://doi.org/10.1007/s12542-009-0038-0.CrossRefGoogle Scholar
Prieto, J., Gross, M., Böhmer, C., and Böhme, M., 2010, Insectivores and bat (Mammalia) from the late Middle Miocene of Gratkorn (Austria): biostratigraphic and ecologic implications: Neues Jahrbuch für Geologie und Paläontologie, v. 258, p. 107119, https://doi.org/10.1127/0077-7749/2010/0088.CrossRefGoogle Scholar
Prieto, J., van den Hoek Ostende, L.W., Hír, J., and Kordos, L., 2015, The Middle Miocene insectivores from Hasznos (Hungary, Nógrád County): Palaeobiodiversity and Palaeoenvironments, v. 95, p. 431451.CrossRefGoogle Scholar
Qiu, Z.D., 1996, Middle Miocene Micromammalian Fauna from Tunggur, Inner Mongolia: Beijing, Science Press, 216 p.Google Scholar
Qiu, Z.D., Wang, X., and Li, Q., 2013, Neogene faunal succession and biochronology of central Nei Mongol (Inner Mongolia), in Wang, X., Flynn, L., and Fortelius, M., Fossil Mammals of Asia: New York, Columbia University Press, v. 1, p. 155186.10.7312/columbia/9780231150125.003.0005CrossRefGoogle Scholar
Radulescu, C., Samson, P., and Știuca, E., 1989, Pliocene (lower Romanian) micromammals in the Dacic Basin: Miscellanea Speologica Romanica, v. 1, p. 313326.Google Scholar
Rümke, C.G., 1974, A new Desmanella species (Talpidae, Insectivora) from the Turolian of Concud and Los Mansuetos (Prov. of Teruel, Spain): Proceedings of the Koninklijke Nederlandse Academie van Wetenschappen, B, v. 77, p. 359374.Google Scholar
Rümke, C.G., 1976, Insectivora from Pikermi and Biodrak (Greece): Proceedings of the Koninklijke Nederlandse Academie van Wetenschappen, B, v. 79, p. 256270.Google Scholar
Rümke, C.G., 1985, A review of fossil and recent Desmaninae (Talpidae, Insectivora): Utrecht Micropaleontological Bulletins, Special Publication, v. 4, 264 p.Google Scholar
Rzebik-Kowalska, B., 2005, Erinaceomorpha and Soricomorpha (Mammalia) from the Miocene of Bełchatów, Poland. IV. Erinaceidae Fischer von Waldheim, 1817 and Talpidae Fischer von Waldheim, 1817: Acta Zoologica Cracoviensia, v. 48, p. 7191.CrossRefGoogle Scholar
Rzebik-Kowalska, B., and Lungu, A., 2009, Insectivore mammals from the Late Miocene of the Republic of Moldova: Acta Zoologica Cracoviensia–Series A: Vertebrata, v. 52, p. 1160, https://doi.org/10.3409/azc.52a_1-2.11-60.Google Scholar
Rzebik-Kowalska, B., and Pawłowski, J., 1994, Ruemkelia (Mammalia, Insectivora, Talpidae) nom. nov. for Dibolia Rümke, 1985 (nec Latreille, 1829): Acta Zoologica Cracoviensia, v. 37, p. 7576.Google Scholar
Rzebik-Kowalska, B., and Rekovets, L.I., 2016, New data on Eulipotyphla (Insectivora, Mammalia) from the Late Miocene to the middle Pleistocene of Ukraine: Palaeontologia Electronica, v. 19, https://doi.org/10.26879/573.Google Scholar
Sabol, M., 2005, Middle Miocene assemblage of insectivores from Bonanza site near Devínska Nová Ves (Slovakia): Geologica Carpathica Bratislava, v. 56, p. 433445.Google Scholar
Sabol, M., Joniak., P., Bilgin, M., Bonilla-Salomón, I., Cailleux, F., Čerňanský, A., Malíková, V., Šedivá, M., and Tóth, C., 2021, Updated Miocene mammal biochronology of Slovakia: Geologica Carpathica, v. 72, p. 425443, https://doi.org/10.31577/GeolCarp.72.5.5.CrossRefGoogle Scholar
Sansalone, G., Colangelo, P., Kotsakis, T, Loy, A., Castiglia, R., Bannikova, A.A., Zemlemerova, E.D., and Piras, P., 2018, Influence of evolutionary allometry on rates of morphological evolution and disparity in strictly subterranean moles (Talpinae, Talpidae, Lipotyphla, Mammalia): Journal of Mammalian Evolution, v. 25, p. 114, https://doi.org/10.1007/s10914-016-9370-9.CrossRefGoogle Scholar
Schreuder, A., 1940, A revision of the fossil water moles (Desmaninae): Archives Néerlandaises de Zoologie, v. 7, p. 202333.Google Scholar
Schwermann, A.H., and Thompson, R.S., 2015 Extraordinarily preserved talpids (Mammalia, Lipotyphla) and the evolution of fossoriality: Journal of Vertebrate Paleontology, v. 35, n. e934828, https://doi.org/10.1080/02724634.2014.934828.CrossRefGoogle Scholar
Storch, G., 1978, Die turolische Wirbeltierfauna von Dorn–Dürkheim, Rheinhessen (SW-Deutschland), 2, Mammalia: Insectivora: Senckenbergiana Lethaea, v. 58, p. 421449.Google Scholar
Storch, G., and Dahlmann, T., 2000, Desmanella rietscheli, ein neuer Talpide aus dem Obermiozän von Dorn Dürkheim 1, Rheinhessen (Mammalia, Lipotyphla): Carolinea, v. 58, p. 6569.Google Scholar
Šujan, M., Braucher, R., Kováč, M., Bourlčs, D. L., Rybár, S., Guillou, V., and Hudáčková, N., 2016, Application of the authigenic 10Be/9Be dating method to Late Miocene–Pliocene sequences in the northern Danube Basin (Pannonian Basin System): confirmation of heterochronous evolution of sedimentary environments: Global and Planetary Change, v. 137, p. 3553, https://doi.org/10.1016/j.gloplacha.2015.12.013.CrossRefGoogle Scholar
Tao, D., 2006, Chinese Neogene mammal biochronology: Vertebrata PalAsiatica, v. 44, p. 143163.Google Scholar
Terzea, E., 1980, Deux micromammifères du Pliocène de Roumanie: Travaux de l'Institut de Spéologie “Émile Racovitza”, v. 19, p. 191201.Google Scholar
Thomas, O., 1912, On a collection of small mammals from the Tsinling Mountains, Central China, presented by Mr. G. Fenwick Owen to the National Museum: Annals and Magazine of Natural History, v. 10, p. 395403.CrossRefGoogle Scholar
Topachevski, V.A., 1961, Novyi Pliotsenovykh vid vykhukholi iz predkavkazya [A new water-mole from the Pliocene]: Paleontological Journal, v. 4, p. 131137.Google Scholar
Topachevsky, V., and Pashkov, A., 1983, Superspecific systematics of the genus Desmana (Insectivora, Talpidae): Vestnik Zoologii, v. 3, p. 3945.Google Scholar
Trouessart, E.-L., 1879, Catalogue des Mammifères vivants et fossiles III. Insectivora: Revue et Magasin de Zoologie Pure et Appliquée, ser. 3, v. 7, p. 219285.Google Scholar
van den Hoek Ostende, L.W., 1989, The Talpidae (Insectivora, Mammalia) of Eggingen–Mittelhart (Baden–Württemberg, F.R.G.) with special reference to the ParatalpaDesmanodon lineage: Stuttgarter Beiträge zur Naturkunde B, v. 152, p. 129.Google Scholar
van den Hoek Ostende, L.W., 1997, Insectivore faunas from the Lower Miocene of Anatolia. Part 4: the genus Desmanodon (Talpidae) with the description of a new species from the Lower Miocene of Spain: Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen, v. 100, p. 2765.Google Scholar
van den Hoek Ostende, L.W., and Fejfar, O., 2006, Erinaceidae and Talpidae (Erinaceomorpha, Soricomorpha, Mammalia) from the Lower Miocene of Merkur–Nord (Czech Republic, MN 3): Beiträge zur Paläontologie, v. 30, p. 175203.Google Scholar
van den Hoek Ostende, L.W., and Furió, M., 2005, Spain, in van den Hoek Ostende, L.W., Doukas, C.S., and Reumer, J.W.F., eds, The Fossil Record of the Eurasian Neogene Insectivores (Erinaceomorpha, Soricomorpha, Mammalia), Part I: Scripta Geologica Special Issue 5, p. 149–284.Google Scholar
Vasileiadou, K., and Doukas, C.S., 2022, The fossil record of insectivores (Mammalia: Eulipotyphla) in Greece, in Vlachos, E., ed., Fossil Vertebrates of Greece, Vol. 2: Cham, Springer, p. 3392.CrossRefGoogle Scholar
Villalta, J.F., and Crusafont, M., 1944, Nuevos insectívoros del Mioceno continental del Vallés-Panadés: Notas y Comunicaciones del Instituto Geológico y Minero de España, v. 12, p. 3965.Google Scholar
Waddell, P.J., Okada, N., and Hasegawa, M., 1999, Towards resolving the interordinal relationships of placental mammals: Systematic Biology, v. 48, p. 15.10.1093/sysbio/48.1.1CrossRefGoogle ScholarPubMed
Ziegler, R., 1985, Talpiden (Mammalia, Insectivora) aus dem Orleanium und Astaracium Bayerns: Mitteilungen der Bayerischen Staatssammlung für Paläontologie und Historische Geologie, v. 25, p. 131175.Google Scholar
Ziegler, R., 1990, Talpidae (Insectivora, Mammalia) aus dem Oberoligozän und Untermiozän Süd-Deutschlands: Stuttgarter Beiträge Naturkunde B, v. 167, p 181.Google Scholar
Ziegler, R., 1994, Bisher übersehene Insectivora (Mammalia) aus dem Unter-miozän von Wintershof−West bei Eichstätt (Bayern): Mitteilungen der Bayerischen Staatssammlung für Paläontologie und Historische Geologie, v. 34, p. 291306.Google Scholar
Ziegler, R., 1998, Marsupialia und Insectivora (Mammalia) aus den oberoligozänen Spaltenfüllungen Herrlingen 8 und Herrlingen 9 bei Ulm (Baden–Württemberg): Senckenbergiana Lethaea, v. 77, p. 101143.CrossRefGoogle Scholar
Ziegler, R., 2003, Moles (Talpidae) from the late Middle Miocene of South Germany: Acta Palaeontologica Polonica, v. 48, p. 617648.Google Scholar
Ziegler, R., 2005, The insectivores (Erinaceomorpha and Soricomorpha, Mammalia) from the Late Miocene hominoid locality Rudabánya: Palaeontographia Italica, v. 90, p. 5381.Google Scholar
Ziegler, R., 2006a, Insectivores (Lipotyphla) and bats (Chiroptera) from the Late Miocene of Austria: Annalen des Naturhistorischen Museums in Wien, Serie A für Mineralogie und Petrographie, Geologie und Paläontologie, Anthropologie und Prähistorie, v. 1, p. 93196.Google Scholar
Ziegler, R., 2006b, Miocene insectivores from Austria and Germany—an overview: Beiträge zur Paläontologie, v. 30, p. 481494.Google Scholar