Penguins (Aves: Sphenisciformes) are interesting to both neontologists and palaeontologists (e.g. Davis & Renner 2003). The fossil record of these extremely specialized inhabitants of the Southern Hemisphere extends back to the Palaeocene epoch (Slack et al. 2006). Extinct penguins are known from localities within the range of their modern-day relatives (Fordyce & Jones 1990), and the oldest diverse assemblage comes from the Eocene La Meseta Formation of Seymour Island, Antarctic Peninsula, the only such locality south of the Antarctic Convergence (Myrcha et al. 2002, Jadwiszczak 2006a). Several collections amounting to over three thousand bones (mainly isolated skeletal elements) have been acquired since 1901 from that formation, and 15 penguin species have been erected so far (Jadwiszczak 2006a, table 1, Tambussi et al. 2006). Only ten of them (grouped into six genera) appear to be taxonomically distinct, and their type specimens are tarsometatarsi (Simpson 1971, Myrcha et al. 2002, Jadwiszczak 2006a, 2006b, p. 296). Individuals from six species belonging to four genera most probably were not larger than those of Aptenodytes forsteri G.R. Gray, 1844, the heaviest and tallest extant penguin (Jadwiszczak 2001, table 3). Interestingly, representatives of all ten species may have co-existed in the West Antarctic during the Late Eocene epoch, just prior to the final break-up of Gondwana (Jadwiszczak 2006a). Presented here is an intriguing partial tarsometatarsus of a small-sized penguin from the Late Eocene of Antarctic Peninsula, probably representing a new genus and species.