Hostname: page-component-7479d7b7d-k7p5g Total loading time: 0 Render date: 2024-07-12T11:41:41.324Z Has data issue: false hasContentIssue false

Ceratozetes and Ceratozetoides (Acari: Oribatida: Ceratozetidae) of North America

Published online by Cambridge University Press:  02 April 2012

Valerie M. Behan-Pelletier*
Affiliation:
Invertebrate Biodiversity Program, Research Branch, Agriculture and Agri-Food Canada, K.W. Neatby Building, Ottawa, Ontario, Canada K1A 0C6
Barbara Eamer
Affiliation:
Invertebrate Biodiversity Program, Research Branch, Agriculture and Agri-Food Canada, K.W. Neatby Building, Ottawa, Ontario, Canada K1A 0C6
*
1Corresponding author (e-mail: Valerie.Behan-Pelletier@agr.gc.ca).

Abstract

The oribatid mite genus Ceratozetes Berlese is represented in America north of Mexico by 21 previously described species known from forest, grassland, subarctic and arctic soils and litter, and canopy habitats. The closely related genus Ceratozetoides Shaldybina is represented in America north of Mexico only by C. cisalpinus (Berlese). Three new species of Ceratozetes from North America are described: C. biporosussp. nov. from forest habitats of southeastern North America, C. cyclopeasp. nov. from montane habitats in western North America, and C. pseudomediocrissp. nov. from forest habitats of western North America. A detailed revised diagnosis is given for Ceratozetes, Ceratozetoides, all previously described species of Ceratozetes (C. angustus (Banks), C. borealis Behan-Pelletier, C. cuspidatus Jacot, C. enodis (Ewing), C. fjellbergi Behan-Pelletier, C. gracilis Michael, C. kutchin Behan-Pelletier, C. longispinus Jacot, C. mediocris (Berlese), C. oresbios Behan-Pelletier, C. pacificus Behan-Pelletier, C. parvulus Sellnick, C. spitsbergensis Thor, C. subaquila (Ewing), C. subinconspicuus (Berlese), C. thienemanni Willmann, C. virginicus (Banks), and C. watertonensis Behan-Pelletier), and Ceratozetoides cisalpinus. Ceratozetes figuratus (Ewing) and C. zeteki (Ewing) are considered junior subjective synonyms of C. enodis (Ewing) syn. nov., and C. inupiaq Behan-Pelletier is transferred to Mycobatidae as Cyrtozetes inupiaq (Behan-Pelletier) comb. nov. New distribution records are given for Ceratozetoides cisalpinus and Ceratozetes angustus, C. borealis, C. cuspidatus, C. gracilis, C. mediocris, C. longispinus, C. oresbios, C. pacificus, C. parvulus, C. thienemanni, C. virginicus, and C. watertonensis. A diagnostic key is provided to adults of the Ceratozetes and Ceratozetoides species now known for America north of Mexico. An analysis of the systematic relationships of 23 of these species based on adult characters indicates that Ceratozetes, excluding Ceratozetoides, is paraphyletic. The clade that includes Ceratozetoides cisalpinus and 15 species of Ceratozetes includes most species for which immatures are known. One sister clade includes Ceratozetes cyclopea, C. enodis, and C. fjellbergi and another includes C. kutchin, C. parvulus, and C. thienemanni. The decision whether or not these 6 species should be retained in Ceratozetessensu stricto awaits discovery of their immatures and molecular analysis.

Résumé

Le genre d’oribates Ceratozetes Berlese est répresenté en Amérique du Nord par 21 espèces qui sont trouvées dans les forêts, les prairies, les sols subarctiques et arctiques, et dans les habitats canopée des forêts. Le genre apparenté Ceratozetoides Shaldybina est répresenté en Amérique du Nord par C. cisalpinus (Berlese). Trois nouvelles espèces de Ceratozetes de l’Amérique du Nord sont proposées et décrites: C. biporosussp. nov. de la litière des forêts du sud-est de l’Amérique du Nord, C. cyclopeasp. nov. des habitats montagneux de l’ouest de l’Amérique du Nord et C. pseudomediocrissp. nov. de la litière des forêts de l’ouest de l’Amérique du Nord. Nous révisons les diagnoses pour Ceratozetes et Ceratozetoides et pour les espèces de Ceratozetes (C. angustus (Banks), C. borealis Behan-Pelletier, C. cuspidatus Jacot, C. enodis (Ewing), C. fjellbergi Behan-Pelletier, C. gracilis Michael, C. kutchin Behan-Pelletier, C. longispinus Jacot, C. mediocris (Berlese), C. oresbios Behan-Pelletier, C. pacificus Behan-Pelletier, C. parvulus Sellnick, C. spitsbergensis Thor, C. subaquila (Ewing), C. subinconspicuus (Berlese), C. thienemanni Willmann, C. virginicus (Banks) et C. watertonensis Behan-Pelletier) et Ceratozetoides cisalpinus. Ceratozetes figuratus (Ewing) et C. zeteki (Ewing) sont considérés comme les synonymes subjectif plus récent du C. enodis (Ewing) syn. nov. et Ceratozetes inupiaq Behan-Pelletier est transféré dans Mycobatidae comme Cyrtozetes inupiaq (Behan-Pelletier) comb. nov. Nous donnons des nouvelles répartitions géographiques pour Ceratozetes angustus, C. borealis, C. cuspidatus, C. gracilis, C. mediocris, C. longispinus, C. oresbios, C. pacificus, C. parvulus, C. thienemanni, C. virginicus et C. watertonensis et pour Ceratozetoides cisalpinus. Nous présentons une clé pour les adults des espèces de Ceratozetes et Ceratozetoides de l’Amérique du Nord. Une analyse des relations systématiques de ces espèces, basée sur les adultes, indique que Ceratozetes, à l’exclusion de Ceratozetoides, est paraphylétique. Le clade qui inclut Ceratozetoides cisalpinus et 15 espèces de Ceratozetes inclut la plupart des espèces pour lesquelles des immatures sont connus. La conservation des deux clades sœurs, Ceratozetes cyclopea, C. enodis, et C. fjellbergi, et C. kutchin, C. parvulus et C. thienemanni dans Ceratozetes sensu stricto attend la découverte de leurs immatures et les analyses moléculaires.

Type
Articles
Copyright
Copyright © Entomological Society of Canada 2009

Access options

Get access to the full version of this content by using one of the access options below. (Log in options will check for institutional or personal access. Content may require purchase if you do not have access.)

References

Alberti, G., Norton, R.A., Adis, J., Fernandez, N.A., Franklin, E., Kratzmann, M., Moreno, A.I., Weigmann, G., and Woas, S. 1997. Porose integumental organs of oribatid mites (Acari, Oribatida). 2. Fine structure. Zoologica, Stuttgart, 146: 33114.Google Scholar
André, M. 1925. Contributions à l'étude des Acariens de la faune française: liste d'Oribatidae receuillies aux environs de Paris. Bulletin du Museum nationale d'Histoire Naturelle Paris, 31: 154158.Google Scholar
Arillo, A., Bordel, I., and Subías, L.S. 1988. Los oribátidos (Acari, Oribatida) de la Ciudad Universitaria de Madrid. Boletín de la Real Sociedad Española de Historia Natural, Sección Biológica, 84: 117125.Google Scholar
Balogh, J., and Balogh, P. 1988. Oribatid mites of the Neotropical Region I. Elsevier, New York.Google Scholar
Balogh, J., and Balogh, P. 1992. The oribatid mites genera of the world. Vols. I and II. Hungarian Natural History Museum, Budapest, Hungary.Google Scholar
Banks, N. 1906. New Oribatidae from the United States. Proceedings of the Academy of Natural Sciences of Philadelphia, 58: 490500.Google Scholar
Banks, N. 1947. On some Acarina from North Carolina. Psyche (Cambridge), 54: 110141.CrossRefGoogle ScholarPubMed
Bayartogtokh, B. 1999. New oribatid mites (Acari: Oribatei) of the genera Dissorhina and Ceratozetoides from Mongolia. Acarologia, 40: 93100.Google Scholar
Behan-Pelletier, V.M. 1984. Ceratozetes (Acari: Ceratozetidae) of Canada and Alaska. The Canadian Entomologist, 116: 14491517CrossRefGoogle Scholar
Behan-Pelletier, V.M. 1985. Ceratozetidae of the western North American Arctic. The Canadian Entomologist, 117: 1287–366.CrossRefGoogle Scholar
Behan-Pelletier, V.M. 1986. Ceratozetidae (Acari: Oribatei) of the western North American subarctic. The Canadian Entomologist, 118: 9911057.CrossRefGoogle Scholar
Behan-Pelletier, V.M. 1997. Oribatid mites (Acari: Oribatida) of the Yukon. In Insects of the Yukon. Edited by Danks, H.V. and Downes, J.A.. Biological Survey of Canada (Terrestrial Arthropods), Ottawa, Ontario. pp. 115149.Google Scholar
Behan-Pelletier, V. 2000. Ceratozetidae (Acari: Oribatida) of arboreal habitats. The Canadian Entomologist, 132: 153182.CrossRefGoogle Scholar
Behan-Pelletier, V. M., and Eamer, B. 2003. Zetomimidae (Acari: Oribatida) of North America. In An acarological tribute to David Cook. Edited by Smith, I.M.. Indira Publishing House, West Bloomfield, Michigan. pp. 2156.Google Scholar
Behan-Pelletier, V.M., and Eamer, B. 2004. Diversity of Oribatida in Canada [online]. Available from http://www.cbif.gc.ca/spp_pages/mites/phps/index_e.php [accessed 1 October 2008].Google Scholar
Berlese, A. 1895. Acari, Myriapoda et Scorpiones hucusque in Italia reperta. Vol. 74. Portici, Padua, Italy. pp. 17.Google Scholar
Berlese, A. 1908. Elenco di generi e specie nuovi di Acari. Redia, 5: 115.Google Scholar
Bernini, F. 1971. Notulae Oribatologicae IV. Contributo alla conoscenza degli Oribatei (Acarida) dei Mti. Reatini (Lazio). Lavori della Societa Italiana di Biogeografia, Nuova Serie, 2: 379400.Google Scholar
Bernini, F., Avanzati, A.M., and Bernini, S. 1988. Notulae Oribatologicae XXXVII. Gli Acari Oribatei del Massiccio del Pollino (Italia Meridionale): aspetti faunistici e biogeografici. Lavori della Societa Italiana di Biogeografia, Nuova Serie, 10(1987): 379488.Google Scholar
Bremer, K. 1994. Branch support and tree stability. Cladistics, 10: 295304.CrossRefGoogle Scholar
Caballero, A.I., and Iturrondobeitia, J.C. 2000. Ácaros Oribátidos interesantes des País Vazco: nuevas citas (Acari, Oribatida). Graellsia, 56: 111114.CrossRefGoogle Scholar
Castagnoli, M., and Pegazzano, F. 1985. Catalogue of the Berlese Acaroteca. Instituto Sperimentale per la Zoologia Agraria, Florence, Italy.Google Scholar
Cianciolo, J.M., and Norton, R.A. 2006. The ecological distribution of reproductive mode in oribatid mites, as related to biological complexity. Experimental and Applied Acarology, 40: 125.CrossRefGoogle ScholarPubMed
Coulson, S.J. 2007. Terrestrial and freshwater invertebrate fauna of the High Arctic archipelago of Svalbard. Zootaxa, 1448: 4158.CrossRefGoogle Scholar
Coulson, S.J., Leinaas, H.P., Ims, R.A., and Søvik, G. 2000. Experimental manipulation of the winter surface ice layer: the effects on a High Arctic soil microarthropod community. Ecography, 23: 299306.CrossRefGoogle Scholar
Ewing, H.E. 1909 a. New American Oribatoidea. Journal of the New York Entomological Society, 17: 116136.Google Scholar
Ewing, H.E. 1909 b. The Oribatoidea of Illinois. Bulletin of the Illinois State Laboratory of Natural History, 7: 337389.CrossRefGoogle Scholar
Ewing, H.E. 1909 c. A systematic and biological study of the Acarina of Illinois. Bulletin, University of Illinois Extension, 7: 359472.Google Scholar
Ewing, H.E. 1917. New Acarina. Part II. Description of new species and varieties from Iowa, Missouri, Illinois, Indiana, and Ohio. Bulletin of the American Museum of Natural History, 37: 149172.Google Scholar
Fujikawa, T. 1996. Oribatid mites from Picea glehni forest at Mt. Ashoro Hokkaido (13). A new species of the family Ceratozetidae. Edaphologia, 57: 2130.Google Scholar
Gjelstrup, P., and Solhøy, T. 1994. The oribatid mites (Acari) of Iceland. The Zoology of Iceland. Steenstrupia, 3: 178.Google Scholar
Goloboff, P.A. 1999. NONA. Version 2. Program and documentation. Fundación e Instituto Miguel Lillo, Tucumán, Argentina.Google Scholar
Grandjean, F. 1940. Les poils et les organes sensitifs portes par les pattes et le palpe chez les Oribates. Deuxième partie. Bulletin de la Société Zoologique de France, 45: 3244.Google Scholar
Grandjean, F. 1941. La chaetotaxie comparée des pattes chez les Oribates (1re série). Bulletin de la Société Zoologique de France, 66: 3350.Google Scholar
Grandjean, F. 1951. Observations sur les Oribates (23e série). Bulletin du Museum Nationale d'Histoire Naturelle. 2nd Series, 2: 261268.Google Scholar
Grandjean, F. 1954. Essai de classification des oribates (acariens). Bulletin de la Société Zoologique de France, 78: 421446.Google Scholar
Grandjean, F. 1962. Nouvelles observations sur les Oribates (2e série). Acarologia, 4: 396422.Google Scholar
Grandjean, F. 1963. Concernant Sphaerobates gratus, les Mochlozetidae et les Ceratozetidae (Oribates). Acarologia, 5: 284305.Google Scholar
Hammer, M. 1952. Investigations on the microfauna of northern Canada. Part I. Oribatidae. Acta Arctica, 4: 1108.Google Scholar
Ibarra, E.L., Wallwork, J.A., and Rodriguez, J.G. 1965. Ecological studies of mites found in sheep and cattle pastures. I. Distribution patterns of oribatid mites. Annals of the Entomological Society of America, 58: 153159.CrossRefGoogle ScholarPubMed
Jacot, A.P. 1936. New moss mites, chiefly Midwestern. American Midland Naturalist, 17: 546553.CrossRefGoogle Scholar
Jacot, A.P. 1937. New moss mites, chiefly Midwestern, II. American Midland Naturalist, 18: 237250.CrossRefGoogle Scholar
Jacot, A.P. 1939. New mites from the White Mountains. Occasional Papers of the Boston Society of Natural History, 8: 321332.Google Scholar
Klironomos, J.N., and Kendrick, W.B. 1995. Relationships among microarthropods, fungi, and their environment. Plant and Soil, 170: 183197.CrossRefGoogle Scholar
Krivolutsky, D.A., and Lebedeva, N.V. 2004. Oribatid mites (Oribatei, Acariformes) in bird feathers: non-passerines. Acta Zoologica Lithuanica, 14: 2647.CrossRefGoogle Scholar
Lamoncha, K.L., and Crossley, D.A. 1998. The oribatid diversity along an elevation gradient in a southeastern Appalachian forest. Pedobiologia, 42: 4355.CrossRefGoogle Scholar
Lebedeva, N.V., Lebedev, V.D., and Melekhina, E.N. 2006. New data on the oribatid mite (Oribatei) fauna of Svalbard. Doklady Biological Sciences (English Translation of Doklady Akademii Nauk SSSR), 407: 182186. [Russian version: Doklady Akademii Nauk SSSR, 407(6): 845–849.]Google ScholarPubMed
Lindberg, N., and Bengtsson, J. 2006. Recovery of forest soil fauna diversity and composition after repeated summer droughts. Oikos, 114: 494506.CrossRefGoogle Scholar
Lindo, Z., and Stevenson, S.K. 2007. Diversity and distribution of oribatid mites (Acari: Oribatida) associated with arboreal and terrestrial habitats in Interior Cedar–Hemlock Forests, British Columbia, Canada. Northwest Science, 81: 305315.CrossRefGoogle Scholar
Lindo, Z., and Visser, S. 2004. Forest floor microarthropod abundance and oribatid mite (Acari: Oribatida) composition following partial and clear-cut harvesting in the mixedwood boreal forest. Canadian Journal of Forest Research, 34: 9981006.CrossRefGoogle Scholar
Lindo, Z., and Winchester, N.N. 2006. A comparison of microarthropod assemblages with emphasis on oribatid mites in canopy suspended soils and forest floors associated with ancient western redcedar trees. Pedobiologia, 50: 3141.CrossRefGoogle Scholar
Lindo, Z., Clayton, M., and Behan-Pelletier, V.M. 2008. Systematics and ecology of Anachipteria geminus sp. nov. (Acari: Oribatida: Achipteriidae) from arboreal lichens in western North America. The Canadian Entomologist, 140: 539556.CrossRefGoogle Scholar
Maddison, D.R., and Maddison, W.P. 2003. MacClade 4: analysis of phylogeny and character evolution. Version 4.06. Sinauer Associates, Sunderland, Massachusetts.Google Scholar
Mahunka, S. 1985. The oribatid fauna of the old juniper woodland of Barcs, Hungary (Acari: Oribatida). Vol. 5. Dunántúli Dolgozatok Termés-zettudomanyi sorozat, Pecs, Hungary. pp. 193206.Google Scholar
Mahunka, S. 1996. Oribatida of the Bükk National Park (Acari, Oribatida). In The fauna of the Bükk National Park. Edited by Mahunka, S.. Hungarian National History Museum, Budapest, Hungary. pp. 491532.Google Scholar
Mahunka, S., and Mahunka-Papp, L. 1995. The oribatid species described by Berlese (Acari). Hungarian Natural History Museum, Budapest, Hungary.Google Scholar
Mahunka, S., and Mahunka-Papp, L. 2004. Contribution to the knowledge of the Hungarian oribatid fauna (Acari) I. Acta Zoologica Academiae Scientiarum Hungaricae, 49: 255260.Google Scholar
Marshall, V.G., Reeves, R.M., and Norton, R.A. 1987. Catalogue of the Oribatida of continental United States and Canada. Memoirs of the Entomological Society of Canada No. 139.CrossRefGoogle Scholar
Menke, H.G. 1963. Revision der Ceratozetidae, Ceratozetes peritus Grandjean (Arach., Acari, Oribatei). Senckenbergiana biologica, 44: 141154.Google Scholar
Menke, H.G. 1964. Revision des Ceratozetidae, 2. Ceratozetes gracilis (Michael) (Arach., Acari, Oribatei). Senckenbergiana biologica, 45: 621634.Google Scholar
Menke, H.G. 1966. Revision der Ceratozetidae, 4. Ceratozetes mediocris Berlese (Arach., Acari, Oribatei). Senckenbergiana biologica, 47: 371378.Google Scholar
Menke, H.G. 1967. Revision der Ceratozetidae, 5. Ceratozetes thienemanni Willmann (Arach., Acari, Oribatei). Senckenbergiana biologica, 48: 415419.Google Scholar
Michael, A.D. 1884. British Oribatidae. Vol. I. The Ray Society, London, United Kingdom.CrossRefGoogle Scholar
Mitchell, M.J. 1976. Ceratozetes kananaskis (Acari: Cryptostigmata: Ceratozetidae): a new mite species from Western Canada. The Canadian Entomologist, 108: 577582.CrossRefGoogle Scholar
Mitchell, M.J. 1977 a. Life history strategies of oribatid mites. In Biology of oribatid mites. Edited by Dindal, D.L.. State University of New York, Syracuse, New York. pp. 6569.Google Scholar
Mitchell, M.J. 1977 b. Population dynamic of oribatid mites (Acari: Cryptostigmata) in an aspen woodland soil. Pedobiologia, 17: 305319.CrossRefGoogle Scholar
Mitchell, M.J. 1978. Vertical and horizontal distribution of oribatid mites (Acari: Cryptostigmata) in an aspen woodland soil. Ecology, 59: 516525.CrossRefGoogle Scholar
Niemi, R., Karppinen, E., and Uusitalo, M. 1997. Catalogue of the Oribatida (Acari) of Finland. Acta Zoologica Fennica, 207: 139.Google Scholar
Norton, R.A. 1977. A review of F. Grandjean's system of leg-chaetotaxy in the Oribatei and its application to the Damaeidae. In Biology of oribatid mites. Edited by Dindal, D.L.. State University of New York, Syracuse, New York. pp. 3362.Google Scholar
Norton, R.A., and Alberti, G. 1997. Porose integumental organs of oribatid mites (Acari, Oribatida). 3. Evolutionary and ecological aspects. Zoologica (Stuttgart), 146: 115143.Google Scholar
Norton, R.A., and Behan-Pelletier, V. M. 2009. Oribatida. In A manual of acarology. 3rd ed. Edited by Krantz, G.W. and Walter, D.E.. Texas Tech University Press, Lubbock, Texas. pp. 430564.Google Scholar
Norton, R.A., and Kethley, J.B. 1989. Berlese North American oribatid mites: historical notes, recombinations, synonymies and type designations. Redia, 62: 421499.Google Scholar
Norton, R.A., Alberti, G., Weigmann, G., and Woas, S. 1997. Porose integumental organs of oribatid mites (Acari, Oribatida). 1. Overview of types and distribution. Zoologica (Stuttgart), 146: 131.Google Scholar
Oudemans, A.C. 1900. New list of Dutch Acari, 1st part. Tijdschrift voor Entomologie, 43: 150171.Google Scholar
Oudemans, A.C. 1913. Acari. In Über Arthropoden in Maulwurfsnestern. Edited by Heselhaus, F.. Tijdschrift voor Entomologie, 56: 224–234, 282.Google Scholar
Pavlitshenko, P.G. 1994. A guide to the ceratozetoid mites (Oribatei, Ceratozetoidea) of Ukraine. Nacionalna Akademija Nauk Ukrainy, Kiev, Ukraine. [In Russian with bilingual keys.]Google Scholar
Pérez-Iñigo, C. 1991. Contribución al conocimiento de las especies Españolas del género Ceratozetes Berlese, 1908 (Acari, Oribatei). Graellsia, 47: 716.Google Scholar
Pérez-Iñigo, C. 1993. Acari, Oribatei, Poronota. In Fauna Iberica. Vol. 3. Edited by Ramos, A., Tercedor, J.A., Ros, X. Bellés i, Noguera, J. Gosálbez i, Sierra, A.G., Mayol, E.M., Piera, F.M., Marina, J.S., and González, J.T.. Museo de Ciencias Naturales, Madrid, Spain. pp. 1320.Google Scholar
Rockett, C.L., and Woodring, J.P. 1966. Biological investigations on a new species of Ceratozetes and of Pergalumna (Acarina: Cryptostigmata). Acarologia, 8: 511520.Google Scholar
Schatz, H. 1995. Hornmilben in Trockenrasenböden des Virgentales (Osttirol, Österreich). 2. Teil: Faunistik (Acari, Oribatida). Berichte des naturwissenschaftlich–medizinischen Vereins in Innsbruck, 82: 121144.Google Scholar
Schatz, H. 2006. Catalogue of known oribatid mite species (Acari Oribatida) from the Central American landbridge (first part). Tropical Zoology, 19: 209288.Google Scholar
Sellnick, M. 1922. Eine neue Oribatide und Berichtigungen zu einer meiner Arbeiten. Schriften der physikalisch–ökonomische Gesellschaft Königsberg, 63: 9798.Google Scholar
Sellnick, M. 1929. Formenkreis: Hornmilben, Oribatei. In Die Tierwelt Mitteleuropas. Vol. 3. Part 9. Edited by Brohmer, P.Ehrmann, P., and Ulmer, G.. Quelle und Meyer, Leipzig, Germany. pp. 142.Google Scholar
Sengbusch, H.G. 1957. Checklist of oribatoid mites in the vicinity of Mountain Lake Biological Station, Virginia. Virginia Journal of Science, 8: 128134.Google Scholar
Seniczak, S. 1993. The morphology of juvenile stages of moss mites of the subfamily Sphaerozetinae (Acari, Oribatida). III. Zoologischer Anzeiger, 231: 2538.Google Scholar
Shaldybina, E.S. 1966. Postembryonic development of beetle mites of the superfamily Ceratozetoidea Balogh, 1961, and their systematics. In Abstracts of the First Acarological Conference, Nos. 225 and 226. Akademia Nauk SSSR, Moscow, USSR. [In Russian.] [Abstr.]Google Scholar
Shaldybina, E.S. 1967 a. The postembryonic development of Ceratozetes cisalpinus Berl., 1908 (Oribatei, Ceratozetidae). Wissenschaftliche Berichte der Höheren Schulen, Biologische Wissenschaften, 11: 1721. [In Russian.]Google Scholar
Shaldybina, E.S. 1967 b. The postembryonic development of Ceratozetes mediocris Berl., 1908 (Oribatei, Ceratozetidae). Zoologicheskii Zhurnal, 46: 692700.Google Scholar
Shaldybina, E.S. 1970. New species of oribatid mites of the subfamily Ceratozetinae (Oribatei) from the Soviet Union. Uchenye zapiski Gor'kovskogo Gosudarstvennogo Pedagogicheskogo Instituta, Seriya Biologiya, 114: 2543. [In Russian.]Google Scholar
Shaldybina, E.S. 1974. Systematics of the Ceratozetidae (Oribatei, Ceratozetoidea). Uchenye zapiski Gor'kovskogo Gosudarstvennogo Pedagogicheskogo Instituta Imeni A M Gor'kogo, 140: 3342. [In Russian.]Google Scholar
Shaldybina, E.S. 1975. Ceratozetoidea. In Soilinhabiting mites: Sarcoptiformes. Edited by Ghilarov, M.S.. Izdatel'stwo Nauka, Moscow, USSR. pp. 275319. [In Russian.]Google Scholar
Shaldybina, E.S. 1977. Preimaginal stages of ceratozetoids (Oribatei, Ceratozetoidea). Fauna, sistematika, biologiya i ekologiya gel'mintov, Gorkij, 7689. [In Russian.]Google Scholar
Skubała, P., and Ciosek, M. 1999. Oribatid mites (Acari, Oribatida) colonizing the zink metallurgic dumps. Fragmenta Faunistica, 42: 4155.CrossRefGoogle Scholar
Skubała, P., and Gulvik, M. 2005. Pioneer oribatid mite communities (Acari, Oribatida) in newly exposed natural (glacier foreland) and anthropogenic (post-industrial dump) habitats. Polish Journal of Ecology, 53: 385407.Google Scholar
Sorenson, M.D. 1999. TreeRot. Version 2. Boston University, Boston, Massachusetts.Google Scholar
Starý, J. 1988. Moss mites (Acari: Oribatida) from some peat bogs in Sumava Mts., south Bohemia. Sborník Jihočeského Muzea v Ceských Budĕjovicích Prirodni Vedy, 28: 99107.Google Scholar
Starý, J. 1994. Oribatid mites (Acari: Oribatida) from the Krkonose Mountains. Opera Corcontica, 31: 115123.Google Scholar
Subías, L.S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) de mundo (1758–2002). Graellsia, 60: 3305.CrossRefGoogle Scholar
Subías, L.S. 2008. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) de mundo (excepto fósiles) [online]. Available from http://www.ucm.es/info/zoo/Artropodos/Catalogo.pdf [accessed 19 February 2009].Google Scholar
Swofford, D.L. 2002. PAUP*: phylogenetic analysis using parsimony (*and other methods). Version 4.0b10. Sinauer Associates, Sunderland, Massachusetts.Google Scholar
Thor, S. 1934. Neue Beiträge zur Kenntnis der Invertebraten-Fauna von Svalbard. Zoologischer Anzeiger, 107: 114139.Google Scholar
Travé, J., and Vachon, M. 1975. François Grandjean 1882–1975 (notice biogeographique et bibliographique). Acarologia, 17: 119.Google Scholar
Weigmann, G. 2006. Hornmilben (Oribatida). Vol. 76. Die Tierwelt Deutschlands.Google Scholar
Willmann, C. 1943. Terrestrische Milben aus Schwedisch-Lappland. Archiv fuer Hydrobiologie, 40: 208239.Google Scholar
Woodring, J.P., and Cook, E.F. 1962 a. The biology of Ceratozetes cisalpinus Berlese, Scheloribates laevigatus Koch, and Oppia neerlandica Oudemans (Oribatei), with a description of all stages. Acarologia, 4: 101137.Google Scholar
Woodring, J.P., and Cook, E.F. 1962 b. The internal anatomy, reproductive physiology, and molting process of Ceratozetes cisalpinus (Acarina, Oribatei). Annals of the Entomological Society of America, 55: 164181.CrossRefGoogle Scholar
Woolley, T.A. 1958. Redescriptions of Ewing's oribatid mites, VIII–IX. Transactions of the American Microscopical Society, 77: 258279.CrossRefGoogle Scholar
Zaitsev, A.S., and Krivolutsky, D.A. 2001. Geography of oribatid mite (Acariformes, Oribatida) diversity in North-European Russia. Mitteilungen der AG Bodenmesofauna (Gieβen), 17: 4349.Google Scholar
Žbikowska-Zdun, K., Piksa, K., and Watrak, I. 2006. Diversity of mites (Acari: Oribatida) in selected microhabitats of the Bug River Protected Landscape Area. Biological Letters, 43: 277286.Google Scholar