Hostname: page-component-848d4c4894-5nwft Total loading time: 0 Render date: 2024-05-31T07:26:29.366Z Has data issue: false hasContentIssue false

Turonian marine amniotes from the Bohemian Cretaceous Basin, Czech Republic

Published online by Cambridge University Press:  22 July 2013

BENJAMIN P. KEAR*
Affiliation:
Palaeobiology Programme, Department of Earth Sciences, Uppsala University, Villavägen 16, SE-752 36 Uppsala, Sweden
BORIS EKRT
Affiliation:
Department of Paleontology, National Museum Prague; Václavské námĕstí 68, 115 79 Prague, Czech Republic
JOSEF PROKOP
Affiliation:
Department of Physical Electronics, Faculty of Nuclear Sciences and Physical Engineering, Czech Technical University in Prague, Břehová 7, 115 19, Prague, Czech Republic
GEORGIOS L. GEORGALIS
Affiliation:
School of Chemical Engineering, Aristotle University of Thessaloniki, Thessaloniki 54 124Greece
*
Author for correspondence: benjamin.kear@geo.uu.se

Abstract

Despite being known for over 155 years, the Late Cretaceous marine amniotes of the Bohemian Cretaceous Basin in the Czech Republic have received little recent attention. These fossils are however significant because they record a diverse range of taxa from an incompletely known geological interval: the Turonian. The presently identifiable remains include isolated bones and teeth, together with a few disarticulated skeletons. The most productive stratigraphical unit is the Lower–Middle Turonian Bílá Hora Formation, which has yielded small dermochelyoid sea turtles, a possible polycotylid plesiosaur and elements compatible with the giant predatory pliosauromorph Polyptychodon. A huge protostegid, together with an enigmatic cheloniid-like turtle, Polyptychodon-like dentigerous components, an elasmosaurid and a tethysaurine mosasauroid have also been found in strata corresponding to the Middle–Upper Turonian Jizera Formation and Upper Turonian – Coniacian Teplice Formation. The compositional character of the Bohemian Cretaceous Basin fauna is compatible with coeval assemblages from elsewhere along the peri-Tethyan shelf of Europe, and incorporates the globally terminal Middle–Upper Turonian occurrence of pliosauromorph megacarnivores, which were seemingly replaced by mosasauroids later in the Cretaceous.

Type
Original Articles
Copyright
Copyright © Cambridge University Press 2013 

Access options

Get access to the full version of this content by using one of the access options below. (Log in options will check for institutional or personal access. Content may require purchase if you do not have access.)

References

Adamovič, J., Čurda, J., Manová, M., Müller, V., Rudolský, J., Rýda, K., Sáňka, V., Stárková, I. & Šalanský, K. 2000. Vysvětlivky k souboru geologických a ekologických účelových map přírodních zdrojů v měřítku. 1:50000. List 14–33 Polička. Český Geologický Ústav, Praha, 76 pp.Google Scholar
Albright, L. B. III, Gillette, D. G. & Titus, A. L. 2007 a. Plesiosaurs from the Upper Cretaceous (Cenomanian–Turonian) Tropic Shale of southern Utah, Part 1: new records of the pliosaur Brachauchenius lucasi . Journal of Vertebrate Paleontology 27, 3140.CrossRefGoogle Scholar
Albright, L. B. Iii, Gillette, D. G. & Titus, A. L. 2007 b. Plesiosaurs from the Upper Cretaceous (Cenomanian–Turonian) Tropic Shale of southern Utah, Part 2: Polycotylidae. Journal of Vertebrate Paleontology 27, 4158.Google Scholar
Andrews, C. W. 1913. A Descriptive Catalogue of the Marine Reptiles of the Oxford Clay. Part II. British Museum (Natural History), London, England, 205 pp.Google Scholar
Andrews, C. W. 1922. Description of a new plesiosaur from the Weald Clay of Berwick (Sussex). Quarterly Journal of the Geological Society of London 78, 285–95.CrossRefGoogle Scholar
Antunes, M. T. 1964. O Neocretácico e o Cenozóico do Litoral de Angola. Junta de Investigaçoes do Ultramar, Lisboa, 254 pp.Google Scholar
Augusta, J. & Soukup, J. 1939. Kritické poznámky k Plesiosauridům z české křídy a nový nález jejich zbytků u Třebovic. Věstník Královské České Společnosti Nauk, Třída Matematicko-Přírodovědecká 1939 (6), 127.Google Scholar
Averianov, A. O. 2001. The first find of a dolichosaur (Squamata, Dolichosauridae) in Central Asia. Paleontologicheskii Zhurnal 35, 525–7.Google Scholar
Bardet, N. & Godefroit, P. 1995. Plesiosaurus houzeaui Dollo, 1909 from the Upper Campanian of Ciply (Belgium) and a review of the Upper Cretaceous plesiosaurs from Europe. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Sciences de la Terre 65, 179–86.Google Scholar
Bardet, N., Houssaye, A., Rage, J.-C. & Pereda Suberbiola, X. 2008. The Cenomanian–Turonian (Late Cretaceous) radiation of marine squamates (Reptilia): the role of the Mediterranean Tethys. Bulletin de la Société Géologique de France 179, 605–22.CrossRefGoogle Scholar
Bardet, N. P., Godefroit, P. & Sciau, J. 1999. A new elasmosaurid plesiosaur from the Lower Jurassic of Southern France. Palaeontology 42, 927–52.CrossRefGoogle Scholar
Bardet, N., Pereda Suberbiola, X. & Jalil, N.-E. 2003 a. A new polycotylid plesiosaur from the Late Cretaceous (Turonian) of Morocco. Comptes Rendus Palevol 2, 305–17.Google Scholar
Bardet, N., Pereda Suberbiola, X. & Jalil, N.-E. 2003 b. A new mosasauroid (Squamata) from the Late Cretaceous (Turonian) of Morocco. Comptes Rendus Palevol 2, 607–16.CrossRefGoogle Scholar
Bardet, N., Pereda Suberbiola, X. & Metais, E. 1998. Un lézard varanoïde (Squamata, Mosasauroidea) dans le Crétacé supérieur deTouraine. Géologie de la France 1, 6972.Google Scholar
Bayer, F. 1896. Kritický seznam plazův a ryb českého útvaru křídového. Věstník České Akademie Císaře Františka Josefa pro Vědy, Slovesnost a Umění 5 (4), 286–94.Google Scholar
Bayer, F. 1897. Ještěr Polyptychodon Ow. Nový nález. Věstník Královské České Společnosti Náuk, Třída Mathematicko-Přírodovědecká 1897 (27), 112.Google Scholar
Bayer, F. 1898. Zpráva o studijní cestě do Londýna. Věstník České Akademie Císaře Františka Josefa pro Vědy, Slovesnost a Umění 7 (2), 107–16.Google Scholar
Bayer, F. 1905. Katalog Českých Fossilních Obratlovců. (Fossilia Vertebrata Bohemiae). České Akademie Vědy Slovesnost a Umění, Praha, 102 pp.CrossRefGoogle Scholar
Bayer, F. 1914. Revise našich ještěrů křídových. Rozpravy České Akademie Císaře Františka Josefa pro Vědy, Slovesnost a Umění 23 (43), 17.Google Scholar
Bell, G. L. Jr. 1997. A phylogenetic revision of North American and Adriatic Mosasauroidea. In Ancient Marine Reptiles (eds Callaway, J. M. & Nicholls, E. L.), pp. 293332. Academic Press, San Diego.CrossRefGoogle Scholar
Bell, G. L. Jr & Polcyn, M. J. 2005. Dallasaurus turneri, a new primitive mosasauroid from the middle Turonian of Texas and comments on the phylogeny of Mosasuridae (Squamata). Netherlands Journal of Geosciences 84, 177–94.Google Scholar
Benson, R. B. J., Butler, R. J., Lindgren, J. & Smith, A. S. 2010. Mesozoic marine tetrapod diversity: mass extinctions and temporal heterogeneity in geological megabiases affecting vertebrates. Proceedings of the Royal Society of London, Series B 277, 829–34.Google Scholar
Brown, D. S. 1981. The English Upper Jurassic Plesiosauroidea (Reptilia) and a review of the phylogeny and classification of the Plesiosauria. Bulletin of the British Museum (Natural History), Geology Series 35, 253347.Google Scholar
Brown, D. S. 1993. A taxonomic reappraisal of the families Elasmosauridae and Cryptoclididae (Reptilia, Plesiosauroidea). Revue de Paléobiologie 7, 916.Google Scholar
Buchy, M.-C. 2006. An elasmosaur (Reptilia: Sauropterygia) from the Turonian (Upper Cretaceous) of Morocco. Carolinea 63, 528.Google Scholar
Buchy, M.-C., Métayer, F. & Frey, E. 2005 a. Osteology of Manemergus anguirostris n. gen. et sp., a new plesiosaur (Reptilia, Sauropterygia) from the Upper Cretaceous of Morocco. Palaeontographica A 272, 97120.Google Scholar
Buchy, M.-C., Smith, K. T., Frey, E., Stinnesbeck, W., Gonzales, A. H. G., Ifrim, C., Lopez-Oliva, J. G. & Porras-Muzquiz, H. 2005 b. Annotated catalogue of marine squamates (Reptilia) from the Upper Cretaceous of northeastern Mexico. Netherlands Journal of Geosciences 84, 195205.Google Scholar
Caldwell, M. W. 1999. Description and phylogenetic relationships of a new species of Coniasaurus Owen, 1850 (Squamata). Journal of Vertebrate Paleontology 19, 438–55.CrossRefGoogle Scholar
Caldwell, M. W. 2000. On the aquatic squamate Dolichosaurus longicollis Owen, 1850 (Cenomanian, Upper Cretaceous), and the evolution of elongate necks in Squamates. Journal of Vertebrate Paleontology 20, 720–35.Google Scholar
Caldwell, M. W. & Lee, M. S. Y. 2004. Reevaluation of the Cretaceous marine lizard Acteosaurus crassicostatus Calligaris, 1993. Journal of Paleontology 78, 617–19.Google Scholar
Caldwell, M. W. & Palci, A. 2007. A new basal mosasauroid from the Cenomanian (U. Cretaceous) of Slovenia with a review of mosasauroid phylogeny and evolution. Journal of Vertebrate Paleontology 27, 863–80.Google Scholar
Carpenter, K. 1996. A review of short-necked plesiosaurs from the Cretaceous of the Western Interior, North America. Neues Jahrbuch für Geologie und Paläontologie Abhandlungen 201, 259–87.Google Scholar
Carpenter, K. 1997. Comparative cranial anatomy of two North American plesiosaurs. In Ancient Marine Reptiles (eds Callaway, J. M. & Nicholls, E. L.), pp. 197203. Academic Press, San Diego.Google Scholar
Carpenter, K. 1999. Revision of North American elasmosaurs from the Cretaceous of the Western Interior. Paludicola 2, 148–73.Google Scholar
Carroll, R. L. & Debraga, M. 1992. Aigialosaurs: mid-Cretaceous varanoid lizards. Journal of Vertebrate Paleontology 12, 6686.Google Scholar
Čech, S. 2011. Palaeogeography and stratigraphy of the Bohemian Cretaceous Basin (Czech Republic) – an overview. Geologické výzkumy na Moravě a ve Slezsku, 2011/1 18, 1821.Google Scholar
Čech, S., Hradecká, L., Svobodová, M. & Švábenická, L. 2005. Cenomanian and Cenomanian–Turonian boundary in the southern part of the Bohemian Cretaceous Basin, Czech Republic. Bulletin of Geosciences 80, 321–54.Google Scholar
Čech, S., Klein, V., Kříž, J. & Valečka, J. 1980. Revision of the Upper Cretaceous stratigraphy of the Bohemian Cretaceous Basin. Věstník Ústředního Ústavu Geologického 55, 277–96.Google Scholar
Collins, J. I. 1970. The chelonian Rhinochelys Seeley from the Upper Cretaceous of England and France. Palaeontology 13, 355–78.Google Scholar
Cope, E. D. 1873. Toxochelys latremis . Proceedings of the Academy of Natural Sciences of Philadelphia 1873, 10.Google Scholar
Debraga, M. & Carroll, R. L. 1993. The origin of mosasaurs as a model of macroevolutionary patterns and processes. Evolutionary Biology 27, 245322.Google Scholar
Diedrich, C. & Hirayama, R. 2003. Turtle remains (Testudines, Chelonioidea) from the middle Turonian of northwest Germany. Netherlands Journal of Geosciences 82, 161–67.CrossRefGoogle Scholar
Druckenmiller, P. & Russell, A. P. 2008. A phylogeny of Plesiosauria (Sauropterygia) and its bearing on the systematic status of Leptocleidus Andrews, 1922 . Zootaxa 1863, 1120.Google Scholar
Dutchak, A. R. & Caldwell, M. W. 2006. Redescription of Aigialosaurus dalmaticus Kramberger, 1892, a Cenomanian mosasauroid lizard from Hvar Island, Croatia. Canadian Journal of Earth Sciences 43, 1821–34.Google Scholar
Edinger, T. 1934. Anton Fritsch's ‘Grosshirn von Polyptychodon’ ist der Steinkern eines Schildkrötenschädels. Psychiatrische en Neurologische Bladen 38 (3–4), 396404.Google Scholar
Ekrt, B. 2012. Reinterpretace nálezu údajné ještĕrky (‘Lacerta sp.’) v české křídĕ. Zprávy o Geologických Výzkumech v Roce 2011 45, 115–16.Google Scholar
Ekrt, B., Čech, S., Košťák, M., Mazuch, M., Voigt, S. & Wiese, F. 2008. New records of teleosts from the late Turonian (Late Cretaceous) of the Bohemian Cretaceous Basin (Czech Republic). Cretaceous Research 29, 659–73.Google Scholar
Ekrt, B., Košťák, M., Mazuch, M., Valíček, J., Voigt, S. & Wiese, F. 2001. Short note on new records of late Turonian (Upper Cretaceous) marine reptile remains from the Úpohlavy quarry (NW Bohemia, Czech republic). Bulletin Czech Geological Survey 76, 101–6.Google Scholar
Ekrt, B., Radoň, M. & Dvořák, P. 2012. Znovuobjevení dvou Fričových originálů ‘mořských plazů’ z české křídy. Zprávy o Geologických Výzkumech v Roce 2011 45, 117–19.Google Scholar
Elliott, D. K. 1997. Desmatochelys lowi, a marine turtle from the Upper Cretaceous. In Ancient Marine Reptiles (eds Callaway, J. M. & Nicholls, E. L.), pp. 243–58. Academic Press, San Diego.Google Scholar
Frič, A. 1877 a. Geologické obrazy země české. IV. Útvar křídový. Vesmír 7, 38–9.Google Scholar
Frič, A. 1877 b. Studien im Gebiete der Böhmischen Kreideformation. Palaeontologische Untersuchungen der einzelnen Schichten. II. Die Weissenberger und Malnitzer Schichten. Archiv der Naturwissenschaftlichen Landesdurchforschung von Böhmen 4 (1), 1153.Google Scholar
Frič, A. 1879. Studie v oboru křídového útvaru v Čechách. Palaeontologické prozkoumání jednotlivých vrstev. II. Bělohorské a Malnické vrstvy. Archiv pro Přírodovědecký Výzkum Čech 4 (1), 1142.Google Scholar
Frič, A. 1889 a. Studien im Gebiete der Böhmischen Kreideformation. Palaeontologische Untersuchungen der einzelnen Schichten. IV. Die Teplitzer Schichten. Archiv der Naturwissenschaftlichen Landesdurchforschung von Böhmen 7 (2), 1120.Google Scholar
Frič, A. 1889 b. Studie v oboru křídového útvaru v Čechách. Palaeontologické prozkoumání jednotlivých vrstev. IV. Teplické vrstvy. Archiv pro Přírodovědecký Výzkum Čech 7 (2), 1114.Google Scholar
Fritsch, A. 1878. Die Reptilien und Fische der Böhmischen Kreideformation. Selbstverlag, Prague, 46 pp.Google Scholar
Fritsch, A. 1905 a. Synopsis der Saurier der böhm. Kreideformation. Sitzungsberichte der Königlich Böhmischen Gesellschaft der Wissenschaften, Classe Mathematisch-Naturwissenschaftliche 1905 (8), 17.Google Scholar
Fritsch, A. 1905 b. Vorläufige Notiz über Miscellanea palaeontologica aus Böhmen und America. Sitzungsberichte der Königlich Böhmischen Gesellschaft der Wissenschaften, Classe Mathematisch-Naturwissenschaftliche 21, 13.Google Scholar
Fritsch, A. 1906. Über neue Saurierfunde in der Kreideformation Böhmens. Sitzungsberichte der Königlich Böhmischen Gesellschaft der Wissenschaften, Classe Mathematisch-Naturwissenschaftliche 1906 (33), 16.Google Scholar
Fritsch, A. 1910. Miscellanea palaeontologica. II. Mesozoica. Selbstverlag, Prague, 26 pp.Google Scholar
Fritsch, A. & Bayer, F. 1905. Neue Fische und Reptilien aus der Böhmischen Kreideformation. Selbstverlag, Prague, 34 pp.Google Scholar
Gradstein, F. J., Ogg, J. G., Schmitz, M. D. & Ogg, G. M. 2012. The Geologic Time Scale 2012. Volume 2. Elsevier, Amsterdam, 1144 pp.Google Scholar
Großmann, F. 2007. The taxonomic and phylogenetic position of the Plesiosauroidea from the Lower Jurassic Posidonia Shale of south-west Germany. Palaeontology 50, 545–64.Google Scholar
Havlíček, P., Holásek, O., Hradecká, L., Jinochová, J., Klečák, J., Majer, V., Manová, M., Müller, V., Rudolský, J., Šalanský, K. & Zelinka, Z. 2001. Vysvětlivky k souboru geologických a ekologických účelových map přírodních zdrojů v měřítku 1:50000. List 13–11 Benátky nad Jizerou. Český Geologický Ústav, Praha, 55 pp.Google Scholar
Hirayama, R. 1994. Phylogenetic systematics of chelonioid sea turtles. The Island Arc 3, 270–84.Google Scholar
Hirayama, R. 1997. Distribution and diversity of Cretaceous chelonioids. In Ancient Marine Reptiles (eds Callaway, J. M. & Nicholls, E. L.), pp. 225–41. Academic Press, San Diego.Google Scholar
Hirayama, R. 1998. Oldest known sea turtle. Nature 392, 705–8.Google Scholar
Hooks, G. E. 1998. Systematic revision of the Protostegidae, with a redescription of Calcarichelys gemma Zangerl, 1953 . Journal of Vertebrate Paleontology 18, 8598.Google Scholar
Houssaye, A. 2010. A new aquatic pythonomorph (Reptilia, Squamata) from the Turonian (Late Cretaceous) of France. Comptes Rendus Palevol 9, 3945.Google Scholar
Jacobs, L. L., Mateus, O., Polcyn, M. J., Schulp, A., Antunes, M. T., Morais, M. L. & Tavares, T. S. 2006. The occurrence and geological setting of Cretaceous dinosaurs, mosasaurs, plesiosaurs, and turtles from Angola. Journal of the Paleontological Society of Korea 22, 91110.Google Scholar
Jacobs, L. L., Polcyn, M. J., Taylor, L. H. & Ferguson, K. 2005. Sea-surface temperatures and palaeoenvironments of dolichosaurs and early mosasaurs. Netherlands Journal of Geosciences 84, 269–81.Google Scholar
Jahn, J. J. 1904. Einige neue Fossilienfundorte in der ostböhmischen Kreideformation. Jahrbuch der Kaiserlich-Königlichen Geologischen Reichsanstalt 54, 7590.Google Scholar
Joyce, W. G. 2007. Phylogenetic relationships of Mesozoic turtles. Bulletin of the Peabody Museum of Natural History 48, 3102.Google Scholar
Karl, H.-V. 2002. Übersicht über die fossilen marinen Schildkrötenfamilien Zentraleuropas (Reptilia, Testudines). Mauritiana (Altenburg) 18, 171202.Google Scholar
Karl, H.-V., Nyhuis, C. J. & Schöllmann, L. 2012. The first shell remains of Rhinochelys (Owen, 1851) from the Upper Cretaceous of NW-Germany (Testudines: Protostegidae). Studia Palaeocheloniologica 4, 143–51.Google Scholar
Kear, B. P. 2002. Reassessment of the Early Cretaceous plesiosaur Cimoliasaurus maccoyi Etheridge, 1904 (Reptilia: Sauropterygia) from White Cliffs, New South Wales. Australian Journal of Zoology 50, 671–85.Google Scholar
Kear, B. P. 2005 a. A new elasmosaurid plesiosaur from the Lower Cretaceous of Queensland, Australia. Journal of Vertebrate Paleontology 25, 792805.Google Scholar
Kear, B. P. 2005 b. Marine reptiles from the Lower Cretaceous (Aptian) deposits of White Cliffs, southeastern Australia: implications of a high-latitude cold water assemblage. Cretaceous Research 26, 769–82.Google Scholar
Kear, B. P. & Barrett, P. M. 2011. Reassessment of the Lower Cretaceous (Barremian) pliosauroid Leptocleidus superstes Andrews, 1922 and other plesiosaur remains from the non-marine Wealden succession of southern England. Zoological Journal of the Linnean Society 161, 663–91.Google Scholar
Kear, B. P. & Lee, M. S. Y. 2006. A primitive protostegid from Australia and early sea turtle evolution. Biology Letters 2, 116–19.Google Scholar
Kear, B. P., Long, J. A. & Martin, J. E. 2005. A review of Australian mosasaur occurrences. Netherlands Journal of Geosciences 84, 307–13.Google Scholar
Kear, B. P., Schroeder, N. I. & Lee, M. S. Y. 2006. An archaic crested plesiosaur in opal from the Lower Cretaceous high latitude deposits of Australia. Biology Letters 2, 615–19.Google Scholar
Ketchum, H. F. & Benson, R. B. J. 2010. Global interrelationships of Plesiosauria (Reptilia, Sauropterygia) and the pivotal role of taxon sampling in determining the outcome of phylogenetic analyses. Biological Reviews 85, 361–92.Google Scholar
Ketchum, H. F. & Benson, R. B. J. 2011. The cranial anatomy and taxonomy of Peloneustes philarchus (Sauropterygia, Pliosauridae) from the Peterborough Member (Callovian, middle Jurassic) of the United Kingdom. Palaeontology 54, 639–65.CrossRefGoogle Scholar
Kollmann, H. A., Peza, L. H. & Čech, S. 1998. Upper Cretaceous Nerineacea of the Bohemian Basin (Czech Republic) and the Saxonian Basin (Germany) and their significance for Tethyan environments. Abhandlungen des staatlichen Museums für Mineralogie und Geologie zu Dresden 43–44, 151–72.Google Scholar
Košťák, M., Čech, S., Ekrt, B., Mazuch, M., Wiese, F., Voigt, S. & Wood, C. J. 2004. Belemnites of the Bohemian Cretaceous Basin in a global context. Acta Geologica Polonica 54, 511–33.Google Scholar
Kubo, T., Mitchell, M. T. & Henderson, D. M. 2012. Albertonectes vanderveldei, a new elasmosaur (Reptilia, Sauropterygia) from the Upper Cretaceous of Alberta. Journal of Vertebrate Paleontology 32, 557–72.Google Scholar
Laube, G. 1896. Pygmaeochelys Michelobana, ein neuer Schildkrötenrest aus dem böhmischen Turon. Sitzungsberichte des Deutschen Naturwissenschaftlich-medicinischen Vereines für Böhmen 16, 2332.Google Scholar
Laurin, J. & Uličný, D. 2004. Controls on a shallow-water hemipelagic carbonate system adjacent to a siliciclastic margin: example from late Turonian of Central Europe. Journal of Sedimentary Research 74, 697717.CrossRefGoogle Scholar
Lee, M. S. Y. & Caldwell, M. W. 2000. Adriosaurus and the affinities of mosasaurs, dolichosaurs, and snakes. Journal of Paleontology 74, 915–37.Google Scholar
Lehman, T. M. & Tomlinson, S. L. 2004. Terlinguachelys fischbecki, a new genus and species of sea turtle (Chelonioidea: Protostegidae) from the Upper Cretaceous of Texas. Journal of Paleontology 78, 1163–78.Google Scholar
Lingham-Soliar, T. 1994. The mosasaur ‘Angolasaurusbocagei (Reptilia: Mosasauridae) from the Turonian of Angola re-interpreted as the earliest member of the genus Platecarpus . Paläontologische Zeitschrift 68, 267–82.Google Scholar
Longman, H. A. 1924. Some Queensland fossil vertebrates. Memoirs of the Queensland Museum 8, 628.Google Scholar
Lydekker, R. 1889 a. On the remains of Eocene and Mesozoic Chelonia and a tooth of (?) Ornithopsis . Quarterly Journal of the Geological Society of London 45, 227–46.CrossRefGoogle Scholar
Lydekker, R. 1889 b. On the remains and affinities of five genera of Mesozoic reptiles. Quarterly Journal of the Geological Society of London 45, 4159.Google Scholar
Makádi, L., Caldwell, M. W. & Ősi, A. 2012. The first freshwater mosasauroid (Upper Cretaceous, Hungary) and a new clade of basal mosasauroids. PLoS ONE 7, e51781. doi:10.1371/journal.pone.0051781 Google Scholar
Mantell, G. 1841. On the remains of turtles, discovered in the Chalk Formation of South-East England. Philosophical Transactions of the Royal Society of London 131, 153–8.Google Scholar
McKean, R. S. 2012. A new species of polycotylid plesiosaur (Reptilia: Sauropterygia) from the lower Turonian of Utah: extending the stratigraphic range of Dolichorhynchops . Cretaceous Research 34, 184–99.Google Scholar
Milner, A. C. 1987. Reptiles. In Fossils of the Chalk (eds Owen, E. & Smith, A. B.), pp. 266–80. Palaeontological Association, London.Google Scholar
Mateus, O., Jacobs, L., Polcyn, M., Schulp, A. S., Vineyard, D., Buta Neto, A. & Telles Antunes, M. 2009. The oldest African eucryptodiran turtle from the Cretaceous of Angola. Acta Palaeontologica Polonica 54, 581–8.Google Scholar
Moody, D. 1993. Cretaceous–Tertiary marine turtles of North West Europe. Revue de Paléobiologie 7, 151–60.Google Scholar
O'keefe, F. R. 2001. A cladistic analysis and taxonomic revision of the Plesiosauria (Reptilia: Sauropterygia). Acta Zoologica Fennica 213, 163.Google Scholar
O'keefe, F. R. 2002. The evolution of plesiosaur and pliosaur morphotypes in the Plesiosauria (Reptilia: Sauropterygia). Paleobiology 28, 101–12.Google Scholar
Owen, R. 1841. Odontography, Part II. Hippolyte Baillière, London, 655 pp.Google Scholar
Owen, R. 1850. Description of the fossil reptiles of the Chalk formations. In The Geology and Fossils of the Tertiary and Cretaceous formations of Sussex (ed. Dixon, E.), pp. 378400. Longman, Brown, Green and Longman, London.Google Scholar
Owen, R. 1851. A monograph on the fossil Reptilia of the Cretaceous formations. Monographs of the Palaeontographical Society 5, 1118.Google Scholar
Páramo-Fonseca, M. E. 2000. Yaguarasaurus columbianus (Reptilia, Mosasauridae), a primitive mosasaur from the Turonian (Upper Cretaceous) of Colombia. Historical Biology 14, 121–31.CrossRefGoogle Scholar
Persson, P. O. 1963. A revision of the classification of the Plesiosauria with a synopsis of the stratigraphical and geographical distribution of the group. Lunds Universitet Årsskrift 65, 123.Google Scholar
Pierce, S. E. & Caldwell, M. W. 2004. Redescription and phylogenetic position of the Adriatic (Upper Cretaceous: Cenomanian) dolichosaur Pontosaurus lesinensis (Kornhuber, 1873). Journal of Vertebrate Paleontology 24, 373–86.Google Scholar
Polcyn, M. J. & Bell, G. L. J. 2005. Russellosaurus coheni n. gen., n. sp., a 92 million-year-old mosasaur from Texas (USA), and the definition of the parafamily Russellosaurina. Netherlands Journal of Geosciences 84, 321–33.Google Scholar
Rage, J.-C. 1989. Le plus ancien lézard varanoïde de France. Bulletin de la Société Études scientifiques d'Anjou 13, 1926.Google Scholar
Reuss, A. E. 1855. Paläontologische miscellen. Denkschriften der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe 10, 7187.Google Scholar
Sachs, S. 2000. Ein pliosauride (Sauropterygia: Plesiosauria) aus der Oberkreide von Anröchte in Westfalen. Geologie und Paläontologie in Westfalen 56, 2534.Google Scholar
Sato, T. 2003. Terminonatator pontiexensis, a new elasmosaur (Reptlia; Sauropterygia) from the Upper Cretaceous of Saskatchewan. Journal of Vertebrate Paleontology 23, 89103.Google Scholar
Sato, T., Hasegawa, Y. & Manabe, M. 2006. A new elasmosaurid plesiosaur from the Upper Cretaceous of Fukushima, Japan. Palaeontology 49, 467–84.Google Scholar
Sato, T., Konishi, T., Hirayama, R. & Caldwell, M. W. 2012. A review of the Upper Cretaceous marine reptiles from Japan. Cretaceous Research 37, 319–40.Google Scholar
Schumacher, B. A. 2008. On the skull of a pliosaur (Plesiosauria; Pliosauridae) from the Upper Cretaceous (early Turonian) of the North American Western Interior. Transactions of the Kansas Academy of Science 111, 203–18.Google Scholar
Schumacher, B. A. 2011. A ‘woollgari-Zone mosasaur’ (Squamata; Mosasauridae) from the Carlile Shale (lower middle Turonian) of central Kansas and the stratigraphic overlap of early mosasaurs and pliosaurid plesiosaurs. Transactions of the Kansas Academy of Science 114, 114.Google Scholar
Schumacher, B. A., Carpenter, K. & Everhart, M. J. 2013. A new Cretaceous pliosaurid (Reptilia, Plesiosauria) from the Carlile Shale (middle Turonian) of Russell County, Kansas. Journal of Vertebrate Paleontology 33, 613–28.Google Scholar
Schumacher, B. A. & Everhart, M. J. 2005. A stratigraphic and taxonomic review of plesiosaurs from the old ‘Fort Benton Group’ of Kansas: a new assessment of old records. Paludicola 5, 3354.Google Scholar
Seeley, H. G. 1874. Note on some generic modifications of the plesiosaurian pectoral arch. Quarterly Journal of the Geological Society of London 30, 436–49.Google Scholar
Smith, K. T. & Buchy, M.-C. 2008. A new aigialosaur (Squamata: Anguimorpha) with soft tissue remains from the Upper Cretaceous of Nuevo León, Mexico. Journal of Vertebrate Paleontology 28, 8594.Google Scholar
Smith, A. S. & Dyke, G. J. 2008. The skull of the giant predatory pliosaur Rhomaleosaurus cramptoni: implications for plesiosaur phylogenetics. Naturwissenschaften 95, 975–80.Google Scholar
Stárková, I. & Opletal, M. 1998. Geologická mapa ČR 1:50000. List 14–33 Polička. Český Geologický Ústav, Praha.Google Scholar
Storrs, G. W. 1991. Anatomy and Relationships of Corosaurus alcovensis (Diapsida: Sauropterygia) and the Triassic Alcova Limestone of Wyoming. Peabody Museum of Natural History, Bulletin 44, 1151.Google Scholar
Storrs, G.W., Arkhangel'skii, M.S. & Efimov, V. 2000. Mesozoic marine reptiles of Russia and other former Soviet republics. In The Age of Dinosaurs in Russia and Mongolia (eds Benton, M. J., Shishkin, M. A., Unwin, D. M. & Kurochkin, E. N.), pp. 187210. Cambridge University Press, Cambridge.Google Scholar
Švábenická, L. 1999. Braarudosphaera-rich sediments in the Turonian of the Bohemian Cretaceous Basin, Czech Republic. Cretaceous Research 20, 773–82.Google Scholar
Svobodová, M., Laurin, J. & Uličný, D. 2002. Palynomorph assemblages in a hemipelagic succession as indicators of transgressive-regressive cycles: example from the upper Turonian of the Bohemian Cretaceous Basin, Czech Republic. In Aspects of Cretaceous Stratigraphy and Palaeobiogeography (ed. Wagreich, M.), pp. 249–68. Verlag der Österreichischen Akademie der Wissenschaften, Vienna.Google Scholar
Tong, H., Hirayama, R., Makhoul, E. & Escuillie, F. 2006. Rhinochelys (Chelonioidea: Protostegidae) from the Late Cretaceous (Cenomanian) of Nammoura, Lebanon. Atti della Societá Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano 147, 113–38.Google Scholar
Uličný, D. 2001. Depositional systems and sequence stratigraphy of coarse-grained deltas in a shallow-marine, strike-slip setting: the Bohemian Cretaceous Basin, Czech Republic. Sedimentology 48, 599628.Google Scholar
Uličný, D., Hladíková, J., Attrep, M. J., Čech, S., Hradecká, L. & Svobodová, M. 1997. Sea-level changes and geochemical anomalies across the Cenomanian–Turonian boundary: Pecínov quarry, Bohemia. Palaeogeography, Palaeoclimatology, Palaeoecology 132, 265–85.Google Scholar
Uličný, D., Laurin, J. & Čech, S. 2009. Controls on clastic sequence geometries in a shallow-marine, transtensional basin: the Bohemian Cretaceous Basin, Czech Republic. Sedimentology 56, 1077–114.Google Scholar
Valeča, J. & Skoček, V. 1991. Late Cretaceous lithoevents in the Bohemian Cretaceous Basin. Cretaceous Research 12, 561–77.Google Scholar
Vincent, P., Bardet, N., Suberbiola, X. P., Bouya, B., Amaghzazm, M. & Meslouh, S. 2011. Zarafasaura oceanis a new elasmosaurid (Reptilia: Sauropterygia) from the Maastrichtian phosphates of Morocco and the palaeobiogeography of latest Cretaceous plesiosaurs. Gondwana Research 19, 1062–73.Google Scholar
Vodrážka, R., Sklenář, J., Čech, S., Laurin, J. & Hradecká, L. 2009. Phosphatic intraclasts in shallow-water hemipelagic strata: a source of palaeoecological, taphonomic and biostratigraphic data (Upper Turonian, Bohemian Cretaceous Basin). Cretaceous Research 30, 204–22.Google Scholar
Voigt, S. & Wiese, F. 2000. Oxygen isotope variations and palaeobiogeographic changes in western and central Europe evidence for late Turonian (Late Cretaceous) climate cooling. Journal of the Geological Society, London 157, 737–44.Google Scholar
VonLoh, J. P. & Bell, , Jr, G. L.. 1998. Fossil reptiles from the Late Cretaceous Greenhorn Formation (late Cenomanian–middle Turonian) of the Black Hills region, South Dakota. Dakoterra 5, 2838.Google Scholar
Welles, S. P. 1943. Elasmosaurid plesiosaurs with a description of new material from California and Colorado. Memoirs of the University of California 13, 125215.Google Scholar
Welles, S. P. 1952. A review of the North American Cretaceous elasmosaurs. University of California Publications in Geological Sciences 29, 44143.Google Scholar
Welles, S. P. 1962. A new species of elasmosaur from the Aptian of Colombia, and a review of the Cretaceous plesiosaurs. University of California Publications in Geological Sciences 44, 196.Google Scholar
Welles, S. P. & Slaughter, R. H. 1963. The first record of the plesiosaurian genus Polyptychodon (Pliosauridae) from the New World. Journal of Paleontology 37, 131–3.Google Scholar
Wieland, G. R. L. 1896. Archelon ischyros: a new gigantic cryptodire testudinate from the Pierre Cretaceous of South Dakota. American Journal of Science 2, 399412.Google Scholar
Wieland, G. R. L. 1909. Revision of the Protostegidae. American Journal of Science 27, 101–30.Google Scholar
Wiese, F., Čech, S., Ekrt, B., Košťák, M., Mazuch, M. & Voigt, S. 2004. The upper Turonian of the Bohemian Cretaceous Basin (Czech Republic) exemplified by the Úpohlavy working quarry: integrated stratigraphy and palaeoceanography of a gateway to the Tethys. Cretaceous Research 25, 329–52.Google Scholar
Wiese, F. & Voigt, S. 2002. Late Turonian (Cretaceous) climate cooling in Europe: faunal response and possible causes. Geobios 35, 6577.Google Scholar
Williston, S. W. 1903. North American plesiosaurs. Part I. Field Columbian Museum Publication 73, Geological Series 2, 179.Google Scholar
Zahálka, Č. 1895. Pásmo IX. Křídového útvaru v okolí Řipu. Nebuželské podolí. Věstník Královské České Společnosti Náuk, Třída Mathematicko-Přírodovědecká 21, 126.Google Scholar
Zahálka, Č. 1896. Palaeontologie křídového útvaru ve Vysočinĕ Řipské a v Polomených Horách. Věstník Královské České Společnosti Náuk, Třída Mathematicko-Přírodovědecká 21, 128.Google Scholar
Zahálka, Č. 1897. Pásmo III. Bělohorské křídového útvaru v poohří. Věstník Královské České Společnosti Náuk, Třída Mathematicko-Přírodovědecká 22, 180.Google Scholar
Zahálka, Č. 1903. Pásmo IX. Křídového útvaru v Pojizeří. Věstník Královské České Společnosti Náuk, Třída Mathematicko-Přírodovĕdecká 32, 1157.Google Scholar
Zangerl, R. 1953 a. The vertebrate fauna of the Selma Formation of Alabama, Part III, the turtles of the family Protostegidae. Fieldiana, Geology Memoirs 3, 59133.Google Scholar
Zangerl, R. 1953 b. The vertebrate fauna of the Selma Formation of Alabama, Part IV, the turtles of the family Toxochelyidae. Fieldiana, Geology Memoirs 3, 137277.Google Scholar
Zangerl, R. 1960. The vertebrate fauna of the Selma Formation of Alabama, Part V, an advanced cheloniid sea turtle. Fieldiana, Geology Memoirs 3, 281312.Google Scholar
Zázvorka, V. 1965. Čelist mosasaurida ze svrchní křídy východních Čech. Časopis Národního Muzea, Oddíl Přírodovědný 134, 217–19.Google Scholar
Ziegler, P. A. 1988. Evolution of the Arctic North Atlantic and the Western Tethys. American Association of Petroleum Geologists Memoir 43, 1198.Google Scholar