Since 1969 archaeological, paleobotanical and ethnographic research in Kagera region, north-western Tanzania (Map 1), have contributed to a deep-time understanding of the ecological history of that region. The primary insight that develops out of this interdisciplinary research is that changes in the ecology of the region can be linked directly to the productive economy and the cultural values of its inhabitants. The development of iron technology and settled agricultural life are the primary components in understanding the environmental changes that arose in this region over the last two millennia or more. This paper provides an overview of the most salient observations derived from this research program. I will begin with a short review of the history of Iron Age settlement, technology and major alterations made to the environment that form the backdrop to the specific reconstructions of forest exploitation gleaned from charcoal excavated from iron-smelting furnaces and from reconstructions of vegetational changes based on palynology.