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Occurrence of Microdictyon from the lower Cambrian Xinji Formation along the southern margin of the North China Platform

Published online by Cambridge University Press:  05 October 2017

Bing Pan ,
Timothy P. Topper ,
Christian B. Skovsted ,
Lanyun Miao  and
Guoxiang Li
Bing Pan
Affiliation:
State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China 〈bpan@nigpas.ac.cn〉, 〈hlmiao@nigpas.ac.cn〉, 〈gxli@nigpas.ac.cn〉 University of Science and Technology of China, Hefei 230026, China
Timothy P. Topper
Affiliation:
Palaeoecosystems Group, Department of Earth Sciences, Durham University, Durham DH1 3LE, United Kingdom 〈timothy.topper@durham.ac.uk〉
Christian B. Skovsted
Affiliation:
Department of Palaeobiology, Swedish Museum of Natural History, Box 50007, SE-104 05 Stockholm, Sweden 〈christian.skovsted@nrm.se〉
Lanyun Miao
Affiliation:
State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China 〈bpan@nigpas.ac.cn〉, 〈hlmiao@nigpas.ac.cn〉, 〈gxli@nigpas.ac.cn〉
Guoxiang Li
Affiliation:
State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China 〈bpan@nigpas.ac.cn〉, 〈hlmiao@nigpas.ac.cn〉, 〈gxli@nigpas.ac.cn〉
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Abstract

Disarticulated net-like plates of the lobopod Microdictyon had a near cosmopolitan distribution from the early to middle Cambrian but are yet to be documented from the North China Platform. Here we report isolated plates of Microdictyon from the lower Cambrian Xinji Formation (Stage 4, Series 2) of the North China Platform, extending the paleogeographic distribution of Microdictyon in the early Cambrian. The plates of Microdictyon from the Xinji Formation are similar to those of other species established on the basis of isolated plates but do bear some new characters, such as mushroom-shaped nodes with a single inclined platform-like apex and an upper surface that displays radial lines. However, the plates documented here are left under open nomenclature due to inadequate knowledge of intraspecific and ontogenetic variation and low specimen numbers. Through comparison of the node shapes of the isolated plates of different Microdictyon species, we consider that low mushroom-shaped nodes could be a primitive and conservative character of Microdictyon while tall mushroom-shaped nodes may be a derived character. Subtle differences in shape and number of node apices may also represent intraspecific or ontogenetic variation.

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Articles
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Journal of Paleontology , Volume 92 , Special Issue 1: Cambrian Explosion , January 2018 , pp. 59 - 70
Copyright
Copyright © 2017, The Paleontological Society 

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References

Bengtson, S., 1991, Oddballs from the Cambrian start to get even: Nature, v. 351, p. 184–185.CrossRefGoogle Scholar
Bengtson, S., Matthews, S.C., and Missarzhevsky, V.V., 1986, The Cambrian netlike fossil Microdictyon , in Hoffman, A., and Nitecki, M.H., eds., Problematic Fossil Taxa: Oxford, UK, Oxford University Press, p. 97–115.Google Scholar
Bengtson, S., Conway Morris, S., Cooper, B.J., Jell, P.A., and Runnegar, B.N., 1990, Early Cambrian fossils from South Australia: Memoirs of the Association of Australasian Palaeontologists, v. 9, p. 332–335.Google Scholar
Bergström, J., and Hou, X., 2001, Cambrian Onychopora or Xenusians: Zoologischer Anzeiger, v. 240, p. 237–245.CrossRefGoogle Scholar
Betts, M.J., Paterson, J.R., Jago, J.B., Jacquet, S.M., Skovsted, C.B., Topper, T.P., and Brock, G.A., 2016, A new lower Cambrian shelly fossil biostratigraphy for South Australia: Gondwana Research, v. 36, p. 163–195.CrossRefGoogle Scholar
Betts, M.J., Paterson, J.R., Jago, J.B., Jacquet, S.M., Skovsted, C.B., Topper, T.P., and Brock, G.A., 2017, Global correlation of the early Cambrian of South Australia: Shelly fauna of the Dailyatia odyssei Zone: Gondwana Research, v. 46, p. 240–279.CrossRefGoogle Scholar
Bureau of Geology and Mineral Resources of Shanxi Province 1989, [Regional Geology of Shanxi Province]: Beijing, Geological Publishing House, p. 116–120 [in Chinese].Google Scholar
Chen, J., Hou, X., and Lu, H., 1989, [Early Cambrian netted scale-bearing wormlike sea animal]: Acta Palaeontologica Sinica, v. 28, p. 1–16 [in Chinese].Google Scholar
Chen, J., Zhou, G., and Ramsköld, L., 1995, The Cambrian lobopodian Microdictyon sinicum: Bulletin of the National Museum of Natural Science, v. 5, p. 1–93.Google Scholar
Conway Morris, S., 1977, A new metazoan from the Cambrian Burgess Shale of British Columbia: Palaeontology, v. 20, p. 623–640.Google Scholar
Conway Morris, S., 1985, The middle Cambrian metazoan Wiwaxia corrugata (Matthew) from the Burgess Shale and Ogygopsis Shale, British Columbia, Canada: Philosophical Transactions of the Royal Society of London, Series B, Biological Sciences, v. 307, p. 507–582.Google Scholar
Conway Morris, S., and Peel, J.S., 1990, Articulated halkieriids from the lower Cambrian of north Greenland: Nature, v. 345, p. 802–805.CrossRefGoogle Scholar
Conway Morris, S., and Peel, J.S., 1995, Articulated halkieriids from the lower Cambrian of North Greenland and their role in early protostome evolution: Philosophical Transactions of the Royal Society of London B, v. 347, p. 305–358.Google Scholar
Dean, W.T., 2005, Trilobites from de Çal tepe formation (Cambrian) near Seydeşehir, central Taurides, southwestern Turkey: Turkish Journal of Earth Sciences, v. 14, no. 1, p. 1–71.Google Scholar
Demidenko, Y.E., 2006, New Cambrian lobopods and chaetognaths of the Siberian Platform: Paleontological Journal, v. 40, no. 3, p. 234–243.CrossRefGoogle Scholar
Dzik, J., 2003, Early Cambrian lobopodian sclerites and associated fossils from Kazakhstan: Palaeontology, v. 46, p. 93–112.CrossRefGoogle Scholar
Dzik, J., and Krumbiegel, G., 1989, The oldest ‘onychophoran’ Xenusion: a link connecting phyla?: Lethaia, v. 22, p. 169–181.CrossRefGoogle Scholar
Esakova, N.V, and Zhegallo, E.A, 1996, [Biostratigraphy and fauna of the lower Cambrian of Mongolia]: Sovmestnaya Rossiisko-Mongolskaya Paleontologicheskaya Ekspeditsiya Trudy, v. 46, p. 1–214 [in Russian].Google Scholar
Feng, W., Qian, Y., and Rong, Z., 1994, [Study of monoplacophora and gastropoda form lower Cambrian Xinji Formation in Yexian, Henan]: Acta Micropalaeontologica Sinica, v. 11, no. 1, p. 1–19 [in Chinese].Google Scholar
Gravestock, D.I., Alexander, E.M., Jago, J.B., Rozanov, A.Y., Zhuravlev, A.Y., et al., 2001, The Cambrian biostratigraphy of the Stansbury Basin, South Australia: Moscow, Transactions of the Palaeontological Institute, v. 282, p. 1–344.Google Scholar
Hao, Y., and Shu, D., 1987, [The oldest known well preserved Pheodaria (Radiolaria) from Southern Shaanxi]: Geoscience, v. 1, no. 3–4, p. 301–310 [in Chinese].Google Scholar
He, T., and Pei, F., 1985, [The discovery of bivalves from the lower Cambrian Xinji Formation in Fangcheng County, Henan Province]: Journal of Chengdu College of Geology, v. 1, p. 61–68 [in Chinese].Google Scholar
He, T., Pei, F., and Fu, G., 1984, [Small shelly fossils from the lower Cambrian Xinji Formation in Fangcheng County, Henan Province]: Acta Palaeontologica Sinica, v. 23, no. 3, p. 350–357 [in Chinese].Google Scholar
Hinz, I., 1987, The lower Cambrian microfauna of Comley and Rushton, Shropshire, England: Palaeontographica Abteilung A, p. 41–100.Google Scholar
Hou, X., and Bergström, J., 1995, Cambrian lobopodians—Ancestors of extant onychophorans?: Zoological Journal of the Linnean Society, v. 114, p. 3–19.CrossRefGoogle Scholar
Hou, X., and Chen, J., 1989, [Early Cambrian arthropod-annelid intermediate sea animal, Luolishania gen. nov. from Chengjiang, Yunnan]: Acta Palaeontologica Sinica, v. 28, no. 2, p. 207–213 [in Chinese].Google Scholar
Hou, X., Ramsköld, L., and Bergström, J., 1991, Composition and preservation of the Chengjiang fauna—A Lower Cambrian soft-bodied biota: Zoologica Scripta, v. 20, p. 395–411.Google Scholar
Hou, X., Aldridge, R.J., Bergström, J., Siveter, D.J., and Feng, X., 2004, The Cambrian Fossils of Chengjiang, China: The Flowering of Early Animal Life: Oxford, UK, Blackwell, p. 1–233.Google Scholar
Hsiang, L., 1963, Trilobita, in Academy of Geological Sciences, Ministry of Geology, ed., Fossil Atlas of Qinling Mountains: Beijing, China Industry Press, 27 p [in Chinese].Google Scholar
Jell, P.A., 1980, Earliest known bivalve on Earth—A new early Cambrian genus from South Australia: Alcheringa, v. 4, p. 233–239.CrossRefGoogle Scholar
Kouchinsky, A., Bengtson, S., Clausen, S., Gubanov, A., Malinky, J.M., and Peel, J.S., 2011, A middle Cambrian fauna of skeletal fossils from the Kuonamka Formation, northern Siberia: Alcheringa, v. 35, p. 123–189.CrossRefGoogle Scholar
Kouchinsky, A., Bengtson, S., Clausen, S., and Vendrasco, M.J., 2015, A lower Cambrian fauna of skeletal fossils from the Emyaksin Formation, northern Siberia: Acta Palaeontologica Polonica, v. 60, no. 2, p. 421–512.Google Scholar
Landing, E., 1988, Lower Cambrian of eastern Massachusetts: Stratigraphy and small shelly fossils: Journal of Paleontology, v. 62, p. 661–695.Google Scholar
Li, G., and Zhu, M., 2001, Discrete sclerites of Microdictyon (lower Cambrian) from the Fucheng section, Nanzheng, South Shaanxi: Acta Palaeontologica Sinica, v. 40, p. 227–235.Google Scholar
Li, G., Zhang, Z., Hua, H., and Yang, H., 2014, Occurrence of the enigmatic bivalved fossil Apistoconcha in the lower Cambrian of southeast Shaanxi, North China platform: Journal of Paleontology, v. 88, no. 2, p. 1–8.CrossRefGoogle Scholar
Li, L., Zhang, X., Yun, H., and Li, G., 2016, New occurrence of Cambroclavus absonus from the lowermost Cambrian of North China and its stratigraphical importance: Alcheringa, v. 40, no. 1, p. 45–52.CrossRefGoogle Scholar
Liu, J., Shu, D., Han, J., and Zhang, Z., 2004, A rare lobopod with well-preserved eyes from the Chengjiang Lagerstätte and its implications for origin of arthropods: Chinese Science Bulletin, v. 49, p. 1063–1071.CrossRefGoogle Scholar
Liu, J., Shu, D., Han, J., Zhang, Z., and Zhang, X., 2006, A large xenusiid lobopod with complex appendages from Chengjiang Lagerstätte (lower Cambrian, China): Acta Palaeontologica Polonica, v. 51, p. 215–222.Google Scholar
Liu, J., Shu, D., Han, J., Zhang, Z., and Zhang, X., 2008, The lobopod Onychodictyon from the lower Cambrian Chengjiang Lagerstätte revisited: Acta Palaeontologica Polonica, v. 53, no. 2, p. 285–292.CrossRefGoogle Scholar
Liu, J., Steiner, M., Dunlop, J.A., Keupp, H., Shu, D., Ou, Q., Han, J., Zhang, Z., and Zhang, X., 2011, An armoured Cambrian lobopodian from China with arthropod-like appendages: Nature, v. 470, p. 526–530.CrossRefGoogle ScholarPubMed
Liu, Q., Ma, L., Zhu, Y., Jin, R., Dai, W., Chen, Y., Fan, X., Zhang, T., and Zhang, Z., 1994, [Lithofacies Palaeogeography and Gypsum Deposits of the Lower Cambrian of North China platform]: Beijing, Geologic Publishing House, 144 [in Chinese].Google Scholar
Liu, Y., 1986, [The lower Cambrian Xinji Formation in Hennan Province]: Henan Geology, v. 4, no. 4, p. 28–35 [in Chinese].Google Scholar
Liu, Y., Wang, J., Zhang, H., and Du, F., 1991, [The Cambrian and Ordovician Systems of Henan Province]: Beijing, Geological Publishing House, 225 p [in Chinese].Google Scholar
Lu, Y., Zhang, W., Zhu, Z., Qian, Y., and Xiang, L., 1965, The Trilobites of China: Beijing, Science Press, 85 p.Google Scholar
Ma, X., Hou, X., and Bergström, J., 2009, Morphology of Luolishania longicruris (lower Cambrian, Chengjiang Lagerstätte, SW China) and the phylogenetic relationships within lobopodians: Arthropod Structure and Development, v. 38, p. 271–291.CrossRefGoogle ScholarPubMed
Matthew, G.F., 1899, Studies on Cambrian faunas: Transactions of the Royal Society of Canada, v. 5, p. 39–66.Google Scholar
Matthews, S., and Missarzhevsky, V., 1975, Small shelly fossils of late Precambrian and early Cambrian age: A review of recent work: Journal of the Geological Society, v. 131, p. 289–304.CrossRefGoogle Scholar
McMenamin, M.A.S., 1984, Paleontology and stratigraphy of lower Cambrian and upper Proterozoic sediments, Caborca region, northwestern Sonora, Mexico [Ph.D. thesis]: Santa Barbara, University of California, 218 p [in Chinese].Google Scholar
McMenamin, M.A.S., and McMenamin, D.L.S., 1990, The Emergence of Animals—The Cambrian Breakthrough: New York, Columbia University Press, 217 p.CrossRefGoogle Scholar
Miao, L., 2014, [Biostratigraphy of the basal Cambrian Xinji Formation and the Houjiashan Formation from the Southern North China Platform] [Master thesis]: Beijing, University of Chinese Academy of Sciences, 127 p [in Chinese].Google Scholar
Miao, L., and Zhu, M., 2014, [Trace fossils from the basal Cambrian Xinji Formation in Southern North China Platform and its chronological significance]: Acta Palaeontologica Sinica, v. 53, no. 3, p. 274–289 [in Chinese].Google Scholar
Missarzhevsky, V.V., 1989, [Oldest skeletal fossils and stratigraphy of Precambrian and Cambrian boundary beds]: Trudy Akademia Nauk SSSR, v. 443, p. 1–236 [in Russian].Google Scholar
Missarzhevsky, V.V., and Mambetov, A.M., 1981, [The stratigraphy and fauna of the Cambrian–Precambrian boundary beds of the Lesser Karatau Range]: Moscow, Trudy Geologischeskogo Instituta Akademia Nauk SSSR, v. 326, p. 1–92 [in Russian].Google Scholar
Pan, B., Miao, L., Yang, H., and Li, G., 2015, [Enigmatic tubular fossil Cupitheca from the lower Cambrian Xinji Formation of Luonan, Shaanxi Province]: Acta Micropalaeontologica Sinica, v. 32, p. 384–395 [in Chinese].Google Scholar
Paterson, J.R., and Brock, G.A., 2007, Early Cambrian trilobites from Angorichina, Flinders Ranges, South Australia, with a new assemblage from the Pararaia bunyerooensis Zone: Journal of Paleontology, v. 81, p. 116–142.CrossRefGoogle Scholar
Pei, F., 1985, [First discovery of Yochelcionella from the lower Cambrian in China and its significance]: Acta Micropalaeontologica Sinica, v. 2, no. 4, p. 395–400 [in Chinese].Google Scholar
Pei, F., and Feng, W., 2005, [Discovery of the molluscan fauna from the lower Cambrian Xinji Formation of Zhuyang, Linbao of Henan]: Journal of Stratigraphy, v. 29, p. 458–461 [in Chinese].Google Scholar
Peng, S., Babcock, L.E., and Cooper, R.A., 2012, The Cambrian Period, in Gradstein, F.M., Ogg, J.G., Schmitz, M.D., and Ogg, G.M., eds., The Geologic Time Scale: Amsrerdam, Elsevier, p. 446.Google Scholar
Pompeckj, J.F., 1927, Ein neues Zeugnis uralten Lebens: Paläontologische Zeitschrift, v. 9, no. 1, p. 287–313 [in German].CrossRefGoogle Scholar
Ramsköld, L., and Chen, J., 1998, Cambrian lobopodians: Morphology and phylogeny, in Edgecombe, G.D., ed., Arthropod Fossils and Phylogeny: New York, Columbia University Press, p. 107–150.Google Scholar
Ramsköld, L., and Hou, X., 1991, New early Cambrian animal and onychophoran affinities of enigmatic metazoans: Nature, v. 351, p. 225–228.CrossRefGoogle Scholar
Sarmiento, G.N., Fernández-Remolar, D., and Göncüoglu, M.C., 2001, Cambrian small shelly fossils from the Cal Tepe Formation, Taurus Mountains: Turkey, Coloquios de Paleontologia, v. 52, p. 117–134.Google Scholar
Shu, D., and Chen, L., 1989, [Discovery of early Cambrian Radiolaria and its significance]: Science in China Series B-Chemistry, v. 32, no. 8, p. 986–994 [in Chinese].Google Scholar
Skovsted, C.B., 2006, Small shelly fauna from the late early Cambrian Bastion and Ella Island Formations, North-East Greenland: Journal of Paleontology, v. 80, p. 1087–1112.CrossRefGoogle Scholar
Skovsted, C.B., Pan, B., Topper, T.P., Betts, M.J., Li, G., and Brock, G.A., 2016, The operculum and mode of life of the lower Cambrian hyolith Cupitheca from South Australia and North China: Palaeogeography, Palaeoclimat-ology, Palaeoecology, v. 443, p. 123–130.CrossRefGoogle Scholar
Smith, M.R., 2014, Ontogeny, morphology and taxonomy of the soft-bodied Cambrian ‘mollusc’: Wiwaxia: Palaeontology, v. 57, p. 215–229.Google Scholar
Smith, M.R., and Ortega-Hernández, J., 2014, Hallucigenia’s onychophoran-like claws and the case for Tactopoda: Nature, v. 514, p. 363–366.CrossRefGoogle ScholarPubMed
Snodgrass, R.E., 1938, Evolution of the Annelida, Onychophora, and Arthropoda: Smithsonian Miscellaneous Collections, v. 97, no. 6, p. 1–159.Google Scholar
Steiner, M., Hu, S., Liu, J., and Keupp, H., 2012, A new species of Hallucigenia from the Cambrian Stage 4 Wulongqing formation of Yunnan (South China) and the structure of sclerites in lobopodians: Bulletin of Geosciences, v. 87, no. 1, 107–124.CrossRefGoogle Scholar
Tong, H., 1989, [A preliminary study on the Microdictyon from the lower Cambrian of Zhenba, South Shaanxi]: Acta Micropalaeontologica Sinica, v. 6, p. 97–101 [in Chinese].Google Scholar
Topper, T.P., Brock, G.A., Skovsted, C.B., and Paterson, J.R., 2009, Shelly fossils from the lower Cambrian Pararaia bunyerooensis Zone, Flinders Ranges, South Australia: Memoirs of the Association of Australasian Palaeontologists, v. 37, p. 199–246.Google Scholar
Topper, T.P., Brock, G.A., Skovsted, C.B., and Paterson, J.R., 2011, Microdictyon plates from the lower Cambrian Ajax Limestone of South Australia: Implications for species taxonomy and diversity: Alcheringa, v. 35, no. 3, p. 427–443.CrossRefGoogle Scholar
Topper, T.P., Skovsted, C.B., Peel, J.S., and Harper, D.A.T., 2013, Moulting in the lobopodian Onychodictyon from the lower Cambrian of Greenland: Lethaia, v. 46, no. 4, p. 490–495.CrossRefGoogle Scholar
Varlamov, A.I., Rozanov, A.Y., Khomentovsky, V.V., Shabanov, Y.Y., Abaimova, G.P., et al., 2008, The Cambrian System of the Siberian Platform, Part 1, The Lena-Aldan Region: Moscow, PIN RAS, 232 p.Google Scholar
Vinther, J., and Nielsen, C., 2005, The early Cambrian Halkieria is a mollusc: Zoologica Scripta, v. 34, p. 81–89.CrossRefGoogle Scholar
Walcott, C.D., 1911, Middle Cambrian annelids: Cambrian geology and paleontology, II: Smithsonian Miscellaneous Collections, v. 57, no. 5, p. 109–142.Google Scholar
Whittington, H.B., 1978, The lobopod animal Aysheaia pedunculata Walcott, middle Cambrian, Burgess Shale, British Columbia: Philosophical Transactions of the Royal Society of London B, v. 284, p. 165–197.Google Scholar
Xiao, L., and Zhou, B., 1984, [Early Cambrian hyolitha from Huainan and Huoqiu county in Anhui province]: Professional Papers of Stratigraphy and Palaeontology, Chinese Academy of Geological Sciences, v. 13, p. 141–151 [in Chinese].Google Scholar
Yan, G., Zhang, E., and Wang, D., 1993, [Evolution of North China-type Cambrian sedimentary environments and trace fossils assemblage in Henan Province]: Sedimentary Facies and Palaeogeograhy, v. 13, no. 3, p. 18–32 [in Chinese].Google Scholar
Yang, B., Steiner, M., and Keupp, H., 2015, Early Cambrian palaeobiogeography of the Zhenba–Fangxian Block (South China): Independent terrane or part of the Yangtze Platform?: Gondwana Research, v. 28, no. 4, p. 1543–1565.CrossRefGoogle Scholar
Yu, W., Xu, J., and Yi, K., 1984, [Discovery of molluscan fauna from lower Cambrian Xinji Formation of Luonan, Shannxi]: Journal of Stratigraphy, v. 8, no. 3, p. 234 [in Chinese].Google Scholar
Yuan, J., Zhu, X., Lin, J., and Zhu, M., 2011, Tentative correlation of Cambrian Series 2 between South China and other continents: Bulletin of Geosciences, v. 86, p. 397–404.CrossRefGoogle Scholar
Zhang, W., and Zhu, Z., 1979, [Notes on some trilobites from lower Cambrian Houjiashan Formation in southern and southwestern parts of North China]: Acta Palaeontologica Sinica, v. 18, no. 6, p. 513–529 [in Chinese].Google Scholar
Zhang, W., Zhu, Z., Yuan, K., Lin, H., Qian, Y., Wu, H., and Yuan, J., 1979, [The boundary of the Cambrian-upper Precambrian in the southern and southwestern parts of North China]: Acta Stratigraphica Sinica, v. 3, no. 1, p. 51–56 [in Chinese].Google Scholar
Zhang, X., and Aldridge, R.J., 2007, Development and diversification of trunk plates of the lower Cambrian lobopodians: Palaeontology, v. 50, p. 401–415.CrossRefGoogle Scholar
Zhao, Y., Yuan, J., Zhu, M., Yang, R., Guo, Q., Qian, Y., Huang, Y., and Pan, Y., 1999, [A progress report on research on the early middle Cambrian Kaili Biota, Guizhou, PRC]: Acta Palaeontologica Sinica, v. 38, supplement, p. 1–14 [in Chinese].Google Scholar
Zhao, Y., Yuan, J., Zhu, M., Yang, R., Guo, Q., Peng, J., and Yang, X., 2002, Progress and significance in research on the early middle Cambrian Kaili Biota, Guizhou Province, China: Progress in Natural Science, v. 12, no. 9, p. 649–654.Google Scholar
Zhao, Y., Zhu, M., Babcock, L.E., Yuan, J., Parsley, R. L., Peng, J., and Wang, Y., 2005, Kaili Biota: A taphonomic window on diversification of metazoans from the basal middle Cambrian: Guizhou, China: Acta Geologica Sinica (English edition), v. 79, no. 6, p. 751–765.Google Scholar
Zhou, B., and Xiao, L., 1984, [Early Cambrian monoplacophorans and gastropods from Huainan and Huoqiu counties (Anhui Provence)]: Professional Papers of Stratigraphy and Palaeontology, v. 13, p. 125–140 [in Chinese].Google Scholar